Asplenium trichomanes |
Asplenium heterochroum |
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doradille chevelue, maidenhair spleenwort |
bicolor spleenwort, varicolor spleenwort |
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Roots | not proliferous. |
not proliferous. |
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Stems | short-creeping, often branched; scales black throughout or with brown borders, lanceolate, 2–5 × 0.2–0.5 mm, margins entire to denticulate. |
erect or ascending, rarely branched; scales black throughout, linear-lanceolate, 2–3 × 0.2–0.4 mm, margin entire, apex attenuate. |
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Leaves | monomorphic. |
monomorphic. |
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Petiole | reddish brown or blackish brown throughout, lustrous, 1–4(–7) cm, 1/6–1/4 length of blade; indument absent or of black, linear-lanceolate or filiform scales at base. |
black or purplish black throughout, lustrous, 0.3–5 cm, 1/4–1/15 length of blade; indument of black filiform scales at base. |
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Blade | linear, 1-pinnate, 3–22 × 0.5–1.5 cm, thin, glabrous or sparsely pubescent; base gradually tapered; apex narrowly acute, not rooting. |
linear or narrowly oblanceolate, 1-pinnate throughout, 6–22(–37) × 1–1.8(–2.3) cm, thin, glabrous; base tapered; apex acute, not rooting. |
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Pinnae | in 15–35 pairs, oblong to oval; medial pinnae 2.5–8 × 2.5–4 mm; base broadly cuneate, with or without low, rounded acroscopic auricle; margins shallowly crenate to serrate or ± entire; apex obtuse. |
in 15–40 pairs, oblong; medial pinnae somewhat asymmetric, oblong, 4–10 × 2–4(–5) mm; base acute, acroscopically enlarged; margins serrate to crenate, lobed; apex obtuse. |
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Veins | free, evident. |
free, evident. |
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Sori | 2–4 pairs per pinna, on both basiscopic and acroscopic sides. |
3–6 pairs per pinna, on both basiscopic and acroscopic sides of pinnae. |
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Spores | 64 per sporangium. |
64 per sporangium. |
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Rachis | reddish brown throughout, lustrous, glabrous or nearly so. |
black throughout, lustrous, glabrous. |
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2n | = 72, 144. |
= 144, 216. |
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Asplenium trichomanes |
Asplenium heterochroum |
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Habitat | Sinkholes, limestone rocks in shady hammocks, masonry | |||||
Elevation | 0–50 m (0–200 ft) | |||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DE; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; QC; Worldwide
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FL; GA; PR; Mexico; West Indies in Cuba; Bermuda; Central America in Belize |
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Discussion | In North America, as in Europe, Asplenium trichomanes consists of diploid and tetraploid cytotypes, treated here as subspecies. Asplenium trichomanes subsp. trichomanes, the diploid, is found on noncalcareous rocks. In the southwestern United States it occurs at high elevations. Asplenium trichomanes subsp. quadrivalens, the tetraploid, grows on calcareous substrates and has a more northern distribution (R. C. Moran 1982). Triploid hybrids are known between the diploids and tetraploids (R. C. Moran 1982; W. H. Wagner Jr. and F. S. Wagner 1966). Subspecies 4 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The tetraploid cytotype (4 x) of Asplenium heterochroum hybridizes with A. resiliens (3 x) to produce A. × heteroresiliens (5 x). It is extremely rare and local, known in northern peninsular Florida from Alachua, Citrus, Columbia, Jackson, and Marion counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | A. muticum | |||||
Name authority | Linnaeus: Sp. Pl. 2: 1080. (1753) | Kunze: Linnaea 9: 67. (1834) | ||||
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