Asplenium trichomanes |
Asplenium rhizophyllum |
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doradille chevelue, maidenhair spleenwort |
doradille ambulante, walking fern, walking spleenwort |
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Roots | not proliferous. |
not proliferous. |
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Stems | short-creeping, often branched; scales black throughout or with brown borders, lanceolate, 2–5 × 0.2–0.5 mm, margins entire to denticulate. |
erect or ascending, usually unbranched; scales dark brown throughout, narrowly deltate, 2–3 × (0.2–)0.5–1 mm, margins entire. |
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Leaves | monomorphic. |
monomorphic but fertile leaves generally larger than sterile leaves. |
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Petiole | reddish brown or blackish brown throughout, lustrous, 1–4(–7) cm, 1/6–1/4 length of blade; indument absent or of black, linear-lanceolate or filiform scales at base. |
reddish brown at base, becoming green distally, dull but sometimes lustrous at base, 0.5–12 cm, 0.1–1.5 times length of blade; indument of dark brown, narrowly deltate scales at base, of minute, club-shaped hairs distally. |
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Blade(s) | linear, 1-pinnate, 3–22 × 0.5–1.5 cm, thin, glabrous or sparsely pubescent; base gradually tapered; apex narrowly acute, not rooting. |
highly variable in size and shape, even on 1 plant, narrowly deltate to linear-lanceolate, simple, 1–30 × 0.5–5 cm, leathery, sparsely pubescent, hairs more numerous abaxially than adaxially; blade base cordate, auriculate, or occasionally hastate, auricles rarely attenuate and radicant; margins entire to sinuate, rarely irregularly incised; apex rounded to very long-attenuate and, if attenuate, generally rooting at tip. |
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Pinnae | in 15–35 pairs, oblong to oval; medial pinnae 2.5–8 × 2.5–4 mm; base broadly cuneate, with or without low, rounded acroscopic auricle; margins shallowly crenate to serrate or ± entire; apex obtuse. |
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Veins | free, evident. |
obscure, anastomosing to form areoles near midrib. |
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Sori | 2–4 pairs per pinna, on both basiscopic and acroscopic sides. |
numerous, scattered somewhat irregularly over blade, often joined at vein junctures. |
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Spores | 64 per sporangium. |
64 per sporangium. |
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Rachis | reddish brown throughout, lustrous, glabrous or nearly so. |
green, dull, nearly glabrous. |
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2n | = 72, 144. |
= 72. |
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Asplenium trichomanes |
Asplenium rhizophyllum |
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Habitat | Shaded, usually moss-covered boulders and ledges, usually on limestone or other basic rocks, but occasionally on sandstone or other acidic rocks, rarely on fallen tree trunks | |||||
Elevation | 0–1000 m (0–3300 ft) | |||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DE; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; QC; Worldwide
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AL; AR; CT; DC; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; TN; VA; VT; WI; WV; ON; QC
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Discussion | In North America, as in Europe, Asplenium trichomanes consists of diploid and tetraploid cytotypes, treated here as subspecies. Asplenium trichomanes subsp. trichomanes, the diploid, is found on noncalcareous rocks. In the southwestern United States it occurs at high elevations. Asplenium trichomanes subsp. quadrivalens, the tetraploid, grows on calcareous substrates and has a more northern distribution (R. C. Moran 1982). Triploid hybrids are known between the diploids and tetraploids (R. C. Moran 1982; W. H. Wagner Jr. and F. S. Wagner 1966). Subspecies 4 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Asplenium rhizophyllum, a diploid species, is morphologically very distinctive within Asplenium and is segregated by many authors, along with its sister species A. ruprechtii Kurata of eastern Asia, into the genus Camptosorus Link. Proliferations arising from leaf tips result in the formation of clonal patches, often dense and extensive, on the mossy boulders and ledges where it typically grows. Naturally occurring sterile hybrids are known with A. platyneuron (these and their fertile allotetraploid derivatives are both referred to A. ebenoides), A. ruta-muraria (A. × inexpectatum E. L. Braun ex C. V. Morton), A. trichomanes subsp. trichomanes [A. × shawneense (R. C. Moran) H. E. Ballard], and A. ebenoides. In addition, the allotetraploid A. pinnatifidum is derived from the hybrid A. montanum × rhizophyllum; the sterile diploid hybrid is unknown. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Camptosorus rhizophyllus | |||||
Name authority | Linnaeus: Sp. Pl. 2: 1080. (1753) | Linnaeus: Sp. Pl. 2: 1078. (1753) | ||||
Web links |
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