Asplenium trichomanes |
Asplenium bradleyi |
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doradille chevelue, maidenhair spleenwort |
Bradley's spleenwort |
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Roots | not proliferous. |
not proliferous. |
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Stems | short-creeping, often branched; scales black throughout or with brown borders, lanceolate, 2–5 × 0.2–0.5 mm, margins entire to denticulate. |
short-creeping to ascending, occasionally branched; scales dark reddish to brown throughout, narrowly deltate, (2–)3–5 × 0.2–0.4 mm, margins entire or shallowly dentate. |
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Leaves | monomorphic. |
monomorphic. |
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Petiole | reddish brown or blackish brown throughout, lustrous, 1–4(–7) cm, 1/6–1/4 length of blade; indument absent or of black, linear-lanceolate or filiform scales at base. |
reddish or purplish brown throughout, lustrous, 1–10(–13) cm, 1/3–3/4 length of blade; indument of brown, narrowly lanceolate scales at base, grading into hairs. |
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Blade | linear, 1-pinnate, 3–22 × 0.5–1.5 cm, thin, glabrous or sparsely pubescent; base gradually tapered; apex narrowly acute, not rooting. |
narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17(–20) × 1–6 cm, thin to moderately thick, sparsely pubescent; base truncate or obtuse; apex acute, not rooting. |
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Pinnae | in 15–35 pairs, oblong to oval; medial pinnae 2.5–8 × 2.5–4 mm; base broadly cuneate, with or without low, rounded acroscopic auricle; margins shallowly crenate to serrate or ± entire; apex obtuse. |
in (3–)5–15(–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate; medial pinnae 6–40 × 3–10 mm; base truncate to obliquely obtuse; margins dentate to denticulate; apex acute or rounded. |
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Veins | free, evident. |
free, barely evident. |
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Sori | 2–4 pairs per pinna, on both basiscopic and acroscopic sides. |
3 to numerous pairs per pinna, on both basiscopic and acroscopic sides. |
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Spores | 64 per sporangium. |
64 per sporangium. |
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Rachis | reddish brown throughout, lustrous, glabrous or nearly so. |
reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent. |
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2n | = 72, 144. |
= 144. |
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Asplenium trichomanes |
Asplenium bradleyi |
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Habitat | Acidic rocks, usually on steep ledges | |||||
Elevation | 0–1000 m (0–3300 ft) | |||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DE; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; QC; Worldwide
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AL; AR; GA; IL; KY; MD; MO; NC; NJ; NY; OH; OK; PA; SC; TN; VA; WV
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Discussion | In North America, as in Europe, Asplenium trichomanes consists of diploid and tetraploid cytotypes, treated here as subspecies. Asplenium trichomanes subsp. trichomanes, the diploid, is found on noncalcareous rocks. In the southwestern United States it occurs at high elevations. Asplenium trichomanes subsp. quadrivalens, the tetraploid, grows on calcareous substrates and has a more northern distribution (R. C. Moran 1982). Triploid hybrids are known between the diploids and tetraploids (R. C. Moran 1982; W. H. Wagner Jr. and F. S. Wagner 1966). Subspecies 4 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Asplenium bradleyi is a morphologically variable species, the allotetraploid derivative of A. montanum × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of A. bradleyi has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, A. bradleyi overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where A. montanum is absent. Sterile hybrids with A. pinnatifidum (A. × gravesii), A. montanum (A. × wherryi D. M. Smith et al.), and A. platyneuron are known from nature. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | A. stotleri | |||||
Name authority | Linnaeus: Sp. Pl. 2: 1080. (1753) | D. C. Eaton: Bull. Torrey Bot. Club 4: 11. (1873) | ||||
Web links |
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