Asplenium rhizophyllum |
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doradille ambulante, walking fern, walking spleenwort |
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Roots | not proliferous. |
Stems | erect or ascending, usually unbranched; scales dark brown throughout, narrowly deltate, 2–3 × (0.2–)0.5–1 mm, margins entire. |
Leaves | monomorphic but fertile leaves generally larger than sterile leaves. |
Petiole | reddish brown at base, becoming green distally, dull but sometimes lustrous at base, 0.5–12 cm, 0.1–1.5 times length of blade; indument of dark brown, narrowly deltate scales at base, of minute, club-shaped hairs distally. |
Blades | highly variable in size and shape, even on 1 plant, narrowly deltate to linear-lanceolate, simple, 1–30 × 0.5–5 cm, leathery, sparsely pubescent, hairs more numerous abaxially than adaxially; blade base cordate, auriculate, or occasionally hastate, auricles rarely attenuate and radicant; margins entire to sinuate, rarely irregularly incised; apex rounded to very long-attenuate and, if attenuate, generally rooting at tip. |
Veins | obscure, anastomosing to form areoles near midrib. |
Sori | numerous, scattered somewhat irregularly over blade, often joined at vein junctures. |
Spores | 64 per sporangium. |
Rachis | green, dull, nearly glabrous. |
2n | = 72. |
Asplenium rhizophyllum |
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Habitat | Shaded, usually moss-covered boulders and ledges, usually on limestone or other basic rocks, but occasionally on sandstone or other acidic rocks, rarely on fallen tree trunks |
Elevation | 0–1000 m (0–3300 ft) |
Distribution |
AL; AR; CT; DC; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; TN; VA; VT; WI; WV; ON; QC
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Discussion | Asplenium rhizophyllum, a diploid species, is morphologically very distinctive within Asplenium and is segregated by many authors, along with its sister species A. ruprechtii Kurata of eastern Asia, into the genus Camptosorus Link. Proliferations arising from leaf tips result in the formation of clonal patches, often dense and extensive, on the mossy boulders and ledges where it typically grows. Naturally occurring sterile hybrids are known with A. platyneuron (these and their fertile allotetraploid derivatives are both referred to A. ebenoides), A. ruta-muraria (A. × inexpectatum E. L. Braun ex C. V. Morton), A. trichomanes subsp. trichomanes [A. × shawneense (R. C. Moran) H. E. Ballard], and A. ebenoides. In addition, the allotetraploid A. pinnatifidum is derived from the hybrid A. montanum × rhizophyllum; the sterile diploid hybrid is unknown. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium |
Sibling taxa | |
Synonyms | Camptosorus rhizophyllus |
Name authority | Linnaeus: Sp. Pl. 2: 1078. (1753) |
Web links |