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Palmer's spleenwort

Morzenti's spleenwort, spleenwort

Roots

not proliferous.

not proliferous.

Stems

short-creeping, unbranched;

scales black with lighter margins, linear-lanceolate, 1.5–3 × 0.1–0.4 mm, margins entire.

erect, rarely branched;

scales black throughout, linear-lanceolate, 1–3.5 × 0.1–0.4 mm, margins entire.

Leaves

monomorphic.

monomorphic.

Petiole

purplish black, lustrous, 0.5–3 cm, 1/3–1/20 length of blade;

indument of black filiform scales at base.

black or purplish black throughout, lustrous, 1–2 cm, 1/5–1/10 length of blade;

indument of black filiform scales at base.

Blade

linear, 1-pinnate throughout, 7–17.5 × 0.9–1.8 cm, thick, glabrous;

base gradually reduced;

apex gradually reduced to whiplike rooting tip.

linear to narrowly oblanceolate, 1-pinnate throughout, 5–15 × 1–2 cm, thick, glabrous;

base gradually tapered, apex acute, not rooting.

Pinnae

in (12–)20–40 pairs, oblong;

medial pinnae 6–9 × 3–4 mm;

base broadly cuneate or auriculate;

margins crenate-serrate;

apex obtuse.

in 12–30 pairs, oblong to quadrangular;

medial pinnae 5–10 × 2–3 mm;

base with slight or prominent acroscopic auricle and often with basiscopic auricle;

margins usually shallowly crenulate, rarely ± entire;

apex acute to obtuse.

Veins

free, obscure.

free, obscure.

Sori

3–7 pairs per pinna, on both basiscopic and acroscopic sides.

1–5 pairs per pinna, on both basiscopic and acroscopic sides.

Spores

64 per sporangium.

32 per sporangium.

Rachis

purplish black throughout, lustrous, glabrous or nearly so.

black or purplish black throughout, lustrous, glabrous or with a few scattered scales.

n

= 2n = 180 (apogamous).

Asplenium palmeri

Asplenium ×heteroresiliens

Habitat Shaded rocky slopes, wet ledges, often in protected places Limestone rocks
Elevation 900–2000(–2750) m (3000–6600(–9000) ft) 0–100 m (0–300 ft)
Distribution
from FNA
AZ; NM; TX; Mexico; Central America in Guatemala; Belize
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; NC; SC
[BONAP county map]
Discussion

V. M. Morzenti (1966) studied the cytology and morphology of Asplenium × heteroresiliens and found it to be a pentaploid (5 x) of hybrid origin between A. resiliens (3 x) and A. heterochroum (4 x). The hybrid is intermediate in morphology between the parents, and it is difficult to separate from them. Misshapen spores mixed with large globose spores can usually be found on the same specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii
Name authority Maxon: Contr. U.S. Natl. Herb. 13: 39. (1909) W.: Amer. Fern J. 56: 12, plate 3. (1966)
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