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Palmer's spleenwort

Bradley's spleenwort

Roots

not proliferous.

not proliferous.

Stems

short-creeping, unbranched;

scales black with lighter margins, linear-lanceolate, 1.5–3 × 0.1–0.4 mm, margins entire.

short-creeping to ascending, occasionally branched;

scales dark reddish to brown throughout, narrowly deltate, (2–)3–5 × 0.2–0.4 mm, margins entire or shallowly dentate.

Leaves

monomorphic.

monomorphic.

Petiole

purplish black, lustrous, 0.5–3 cm, 1/3–1/20 length of blade;

indument of black filiform scales at base.

reddish or purplish brown throughout, lustrous, 1–10(–13) cm, 1/3–3/4 length of blade;

indument of brown, narrowly lanceolate scales at base, grading into hairs.

Blade

linear, 1-pinnate throughout, 7–17.5 × 0.9–1.8 cm, thick, glabrous;

base gradually reduced;

apex gradually reduced to whiplike rooting tip.

narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17(–20) × 1–6 cm, thin to moderately thick, sparsely pubescent;

base truncate or obtuse;

apex acute, not rooting.

Pinnae

in (12–)20–40 pairs, oblong;

medial pinnae 6–9 × 3–4 mm;

base broadly cuneate or auriculate;

margins crenate-serrate;

apex obtuse.

in (3–)5–15(–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate;

medial pinnae 6–40 × 3–10 mm;

base truncate to obliquely obtuse;

margins dentate to denticulate;

apex acute or rounded.

Veins

free, obscure.

free, barely evident.

Sori

3–7 pairs per pinna, on both basiscopic and acroscopic sides.

3 to numerous pairs per pinna, on both basiscopic and acroscopic sides.

Spores

64 per sporangium.

64 per sporangium.

Rachis

purplish black throughout, lustrous, glabrous or nearly so.

reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent.

2n

= 144.

Asplenium palmeri

Asplenium bradleyi

Habitat Shaded rocky slopes, wet ledges, often in protected places Acidic rocks, usually on steep ledges
Elevation 900–2000(–2750) m (3000–6600(–9000) ft) 0–1000 m (0–3300 ft)
Distribution
from FNA
AZ; NM; TX; Mexico; Central America in Guatemala; Belize
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; GA; IL; KY; MD; MO; NC; NJ; NY; OH; OK; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Asplenium bradleyi is a morphologically variable species, the allotetraploid derivative of A. montanum × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of A. bradleyi has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, A. bradleyi overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where A. montanum is absent. Sterile hybrids with A. pinnatifidum (A. × gravesii), A. montanum (A. × wherryi D. M. Smith et al.), and A. platyneuron are known from nature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. stotleri
Name authority Maxon: Contr. U.S. Natl. Herb. 13: 39. (1909) D. C. Eaton: Bull. Torrey Bot. Club 4: 11. (1873)
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