Asplenium montanum |
Asplenium heterochroum |
|
---|---|---|
mountain spleenwort |
bicolor spleenwort, varicolor spleenwort |
|
Roots | proliferous. |
not proliferous. |
Stems | horizontal, often arching upward, unbranched (although clusters of stems often form from root proliferations, giving false appearance of single much-branched stem); scales dark brown throughout, narrowly deltate, 2–4 × 0.2–0.4 mm, margins entire. |
erect or ascending, rarely branched; scales black throughout, linear-lanceolate, 2–3 × 0.2–0.4 mm, margin entire, apex attenuate. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark brown to purplish black, lustrous proximally, fading to green distally, 2–11 cm, 1/2–1 1/2 length of blade; indument of blackish, narrowly lanceolate scales only at very base and of minute hairs. |
black or purplish black throughout, lustrous, 0.3–5 cm, 1/4–1/15 length of blade; indument of black filiform scales at base. |
Blade | deltate to lanceolate, 1–2-pinnate-pinnatifid, 2–11 × 1–7(–10) cm, thick, essentially glabrous; base truncate or obtuse; apex acuminate to acute, not rooting. |
linear or narrowly oblanceolate, 1-pinnate throughout, 6–22(–37) × 1–1.8(–2.3) cm, thin, glabrous; base tapered; apex acute, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; proximal (longest) pinnae 6–35 × 4–20 mm; base obtuse to acute; margins coarsely incised; apex acute to rounded. |
in 15–40 pairs, oblong; medial pinnae somewhat asymmetric, oblong, 4–10 × 2–4(–5) mm; base acute, acroscopically enlarged; margins serrate to crenate, lobed; apex obtuse. |
Veins | free, obscure. |
free, evident. |
Sori | 1–15 per pinna, on both basiscopic and acroscopic sides. |
3–6 pairs per pinna, on both basiscopic and acroscopic sides of pinnae. |
Spores | 64 per sporangium. |
64 per sporangium. |
Rachis | green throughout, dull, sparsely pubescent. |
black throughout, lustrous, glabrous. |
2n | = 72. |
= 144, 216. |
Asplenium montanum |
Asplenium heterochroum |
|
Habitat | Crevices in sandstone or other acidic rocks | Sinkholes, limestone rocks in shady hammocks, masonry |
Elevation | 0–2000 m (0–6600 ft) | 0–50 m (0–200 ft) |
Distribution |
AL; CT; GA; IN; KY; MA; MD; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; WV
|
FL; GA; PR; Mexico; West Indies in Cuba; Bermuda; Central America in Belize |
Discussion | Asplenium montanum occurs principally in the Appalachian region, with outlying localities in the Shawnee Hills of western Kentucky (R. Cranfill 1980) and adjacent Indiana (D. M. Smith 1956). A report of its disjunct occurrence on the northern edge of the Ozarks is based on a single specimen whose label indicates the collection locality near Graham Cave, Montgomery County, Missouri. Efforts by several botanists to relocate the population have failed. Reports of a disjunct station in the upper peninsula of Michigan are doubtful. Asplenium montanum is an ecological specialist. It is typically the sole vascular plant species in the siliceous rock crevices in which it is found. It may occur, however, with two allotetraploid species, A. bradleyi and A. pinnatifidum, which were derived from hybrids of A. montanum with A. platyneuron and A. rhizophyllum, respectively. In addition, A. montanum crosses frequently with A. pinnatifidum producing A. × trudellii and rarely with allotetraploid individuals of A. bradleyi producing A. × wherryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The tetraploid cytotype (4 x) of Asplenium heterochroum hybridizes with A. resiliens (3 x) to produce A. × heteroresiliens (5 x). It is extremely rare and local, known in northern peninsular Florida from Alachua, Citrus, Columbia, Jackson, and Marion counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. muticum | |
Name authority | Willdenow: Sp. Pl. 5(1): 342. (1810) | Kunze: Linnaea 9: 67. (1834) |
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