Asplenium adiantum-nigrum |
Asplenium ×heteroresiliens |
|
---|---|---|
black spleenwort |
Morzenti's spleenwort, spleenwort |
|
Roots | not proliferous. |
not proliferous. |
Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
erect, rarely branched; scales black throughout, linear-lanceolate, 1–3.5 × 0.1–0.4 mm, margins entire. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
black or purplish black throughout, lustrous, 1–2 cm, 1/5–1/10 length of blade; indument of black filiform scales at base. |
Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
linear to narrowly oblanceolate, 1-pinnate throughout, 5–15 × 1–2 cm, thick, glabrous; base gradually tapered, apex acute, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
in 12–30 pairs, oblong to quadrangular; medial pinnae 5–10 × 2–3 mm; base with slight or prominent acroscopic auricle and often with basiscopic auricle; margins usually shallowly crenulate, rarely ± entire; apex acute to obtuse. |
Veins | free, evident. |
free, obscure. |
Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
1–5 pairs per pinna, on both basiscopic and acroscopic sides. |
Spores | 64 per sporangium. |
32 per sporangium. |
Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
black or purplish black throughout, lustrous, glabrous or with a few scattered scales. |
n | = 2n = 180 (apogamous). |
|
2n | = 144. |
|
Asplenium adiantum-nigrum |
Asplenium ×heteroresiliens |
|
Habitat | Cliffs | Limestone rocks |
Elevation | 1675–2300 m (5500–7500 ft) | 0–100 m (0–300 ft) |
Distribution |
AZ; CO; UT; Eurasia; Africa |
FL; NC; SC |
Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
V. M. Morzenti (1966) studied the cytology and morphology of Asplenium × heteroresiliens and found it to be a pentaploid (5 x) of hybrid origin between A. resiliens (3 x) and A. heterochroum (4 x). The hybrid is intermediate in morphology between the parents, and it is difficult to separate from them. Misshapen spores mixed with large globose spores can usually be found on the same specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | |
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | W.: Amer. Fern J. 56: 12, plate 3. (1966) |
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