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black spleenwort

Biscayne spleenwort

Roots

not proliferous.

proliferous.

Stems

ascending or short-creeping, infrequently branched;

scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate.

erect, not branched;

scales blackish throughout, linear-deltate, sparse, 1–1.3 × 0.1–2.4 mm, margins entire.

Leaves

monomorphic.

nearly monomorphic.

Petiole

dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade;

indument of black filiform scales and minute hairs.

green, becoming blackish in older leaves, 1–5(–12) cm, 1/8–1/3 length of blade;

indument of black, narrowly lanceolate scales.

Blade

deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute;

base truncate;

apex acute to acuminate, not rooting.

dull, linear, 2-pinnate, (4–)12–22 × 1–3.5 cm, papery, glabrous;

base slightly tapered;

apex narrowing gradually, not rooting.

Pinnae

in 4–10 pairs, deltate to lanceolate;

most proximal (largest) pinnae 1.5–4 × 1–2.5 cm;

base obliquely obtuse;

segment margins coarsely incised;

apex acute.

in 9–20 pairs, oblong, 0.5–2 × 0.4–1.3 cm;

apex blunt to truncate.

Veins

free, evident.

free, evident.

Sori

1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides.

1–2 per segment, on both basiscopic and acroscopic sides.

Spores

64 per sporangium.

abortive.

Rachis

greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent.

mostly blackish, green distally and in juvenile leaves, shiny, sparsely scaly.

Pinnules

of 1–2 segments;

segments linear-oblong, 3–6 × 1–3 mm;

margins dentate;

apex notched, pointed, rounded, or blunt.

2n

= 144.

= ca. 180.

Asplenium adiantum-nigrum

Asplenium ×biscayneanum

Habitat Cliffs Hammocks, on limestone rock faces
Elevation 1675–2300 m (5500–7500 ft) 0–20 m (0–100 ft)
Distribution
from FNA
AZ; CO; UT; Eurasia; Africa
[BONAP county map]
from FNA
FL
Discussion

Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Asplenium × biscayneanum is the hybrid of A. trichomanes-dentatum and A. verecundum, with which it occurs. This peculiar spleenwort may most readily be separated from A. trichomanes-dentatum by its deeply cut pinnae, and from A. verecundum by being only 2-pinnate and having long petioles. Chromosome pairing in A. × biscayneanum is irregular. Judging from herbarium collections, it shows considerable hybrid vigor. All the collections are from Dade County, Florida, where it may now be extirpated.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. andrewsii, A. chihuahuense, A. dubiosum A. rhizophyllum var. biscayneanum
Name authority Linnaeus: Sp. Pl. 2: 1081. (1753) (D. C. Eaton) A. A. Eaton: Fern Bull. 12: 45. (1904)
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