Asplenium adiantum-nigrum |
Asplenium vespertinum |
|
---|---|---|
black spleenwort |
western spleenwort |
|
Roots | not proliferous. |
not proliferous. |
Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
short-creeping, rarely branched; scales black with lighter margins, linear-lanceolate, 2–3 × 0.2–0.4 mm, margins entire, or denticulate distally. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
reddish brown or purplish black throughout, lustrous, 2–5 cm, 1/4–1/6 length of blade, sometimes curved like a J at base; indument of blackish brown, filiform scales at base. |
Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
linear, 1-pinnate throughout, 5–15(–28) × 1–2.5 cm, thick, glabrous; base gradually tapered; apex acute, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
15–30 pairs, oblong; medial pinnae 5–10 × 2–3 mm; base rounded to cuneate; margins lobed or serrate; apex obtuse. |
Veins | free, evident. |
free, obscure. |
Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
2–6 pairs per pinna, on both basiscopic and acroscopic sides. |
Spores | 64 per sporangium. |
64 per sporangium. |
Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
reddish brown or purplish black throughout, shiny, glabrous. |
2n | = 144. |
|
Asplenium adiantum-nigrum |
Asplenium vespertinum |
|
Habitat | Cliffs | Moist, shaded canyon walls and at base of overhanging rocks |
Elevation | 1675–2300 m (5500–7500 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AZ; CO; UT; Eurasia; Africa |
CA; Mexico in Baja California
|
Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | A. trichomanes var. vespertinum |
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | Maxon: Bull. Torrey Bot. Club 27: 197. (1900) |
Web links |