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black spleenwort

western spleenwort

Roots

not proliferous.

not proliferous.

Stems

ascending or short-creeping, infrequently branched;

scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate.

short-creeping, rarely branched;

scales black with lighter margins, linear-lanceolate, 2–3 × 0.2–0.4 mm, margins entire, or denticulate distally.

Leaves

monomorphic.

monomorphic.

Petiole

dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade;

indument of black filiform scales and minute hairs.

reddish brown or purplish black throughout, lustrous, 2–5 cm, 1/4–1/6 length of blade, sometimes curved like a J at base;

indument of blackish brown, filiform scales at base.

Blade

deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute;

base truncate;

apex acute to acuminate, not rooting.

linear, 1-pinnate throughout, 5–15(–28) × 1–2.5 cm, thick, glabrous;

base gradually tapered;

apex acute, not rooting.

Pinnae

in 4–10 pairs, deltate to lanceolate;

most proximal (largest) pinnae 1.5–4 × 1–2.5 cm;

base obliquely obtuse;

segment margins coarsely incised;

apex acute.

15–30 pairs, oblong;

medial pinnae 5–10 × 2–3 mm;

base rounded to cuneate;

margins lobed or serrate;

apex obtuse.

Veins

free, evident.

free, obscure.

Sori

1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides.

2–6 pairs per pinna, on both basiscopic and acroscopic sides.

Spores

64 per sporangium.

64 per sporangium.

Rachis

greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent.

reddish brown or purplish black throughout, shiny, glabrous.

2n

= 144.

Asplenium adiantum-nigrum

Asplenium vespertinum

Habitat Cliffs Moist, shaded canyon walls and at base of overhanging rocks
Elevation 1675–2300 m (5500–7500 ft) 0–1000 m (0–3300 ft)
Distribution
from FNA
AZ; CO; UT; Eurasia; Africa
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. andrewsii, A. chihuahuense, A. dubiosum A. trichomanes var. vespertinum
Name authority Linnaeus: Sp. Pl. 2: 1081. (1753) Maxon: Bull. Torrey Bot. Club 27: 197. (1900)
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