Asplenium adiantum-nigrum |
Asplenium verecundum |
|
---|---|---|
black spleenwort |
limestone spleenwort |
|
Roots | not proliferous. |
proliferous. |
Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
erect, unbranched; scales blackish throughout, linear to narrowly triangular, 0.9–1.3 × 0.1–0.4 mm, margins entire. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
brownish black, 0.5–3(–5) cm, 1/9–1/7 length of blade; indument of blackish, linear-lanceolate scales at base. |
Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
narrowly lanceolate, 2–3-pinnate, (4–)10–20(–)30 × 1–3(–5) cm, thin, glabrous; base tapered somewhat; apex formed by gradual reduction, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
in (8–)12–16(–22) pairs, oblong; medial pinnae 0.5–2.5 × 3–10 mm; base cuneate; apex obtuse. |
Veins | free, evident. |
free, evident, 1 per segment. |
Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
1 per segment. |
Spores | 64 per sporangium. |
64 per sporangium. |
Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
blackish brown, dull, essentially glabrous. |
Pinnules | of 1–5 segments; segments oblanceolate, 2–3 × 1–2 mm, entire; apex mostly round. |
|
2n | = 144. |
= 144. |
Asplenium adiantum-nigrum |
Asplenium verecundum |
|
Habitat | Cliffs | Limestone outcrops in grottoes, on cliffs, and on boulders in shaded woods |
Elevation | 1675–2300 m (5500–7500 ft) | 0–50 m (0–200 ft) |
Distribution |
AZ; CO; UT; Eurasia; Africa |
FL |
Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
This delicate spleenwort, Asplenium verecundum, occurs in both southern and central peninsular Florida where it is very local. Sterile hybrids are known to result from crossing with A. trichomanes-dentatum (A. × biscayneanum) and A. abscissum (A. × curtissii). Asplenium verecundum may be a variety or cytotype of the similar A. myriophyllum (Swartz) C. Presl of the West Indies (D. B. Lellinger 1985). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | A. scalifolium, A. suare, A. subtile |
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | Chapman ex L. Underwood: Bull. Torrey Bot. Club 33: 193. (1906) |
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