Asplenium adiantum-nigrum |
Asplenium trichomanes |
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black spleenwort |
doradille chevelue, maidenhair spleenwort |
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Roots | not proliferous. |
not proliferous. |
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Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
short-creeping, often branched; scales black throughout or with brown borders, lanceolate, 2–5 × 0.2–0.5 mm, margins entire to denticulate. |
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Leaves | monomorphic. |
monomorphic. |
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Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
reddish brown or blackish brown throughout, lustrous, 1–4(–7) cm, 1/6–1/4 length of blade; indument absent or of black, linear-lanceolate or filiform scales at base. |
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Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
linear, 1-pinnate, 3–22 × 0.5–1.5 cm, thin, glabrous or sparsely pubescent; base gradually tapered; apex narrowly acute, not rooting. |
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Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
in 15–35 pairs, oblong to oval; medial pinnae 2.5–8 × 2.5–4 mm; base broadly cuneate, with or without low, rounded acroscopic auricle; margins shallowly crenate to serrate or ± entire; apex obtuse. |
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Veins | free, evident. |
free, evident. |
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Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
2–4 pairs per pinna, on both basiscopic and acroscopic sides. |
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Spores | 64 per sporangium. |
64 per sporangium. |
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Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
reddish brown throughout, lustrous, glabrous or nearly so. |
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2n | = 144. |
= 72, 144. |
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Asplenium adiantum-nigrum |
Asplenium trichomanes |
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Habitat | Cliffs | |||||
Elevation | 1675–2300 m (5500–7500 ft) | |||||
Distribution |
AZ; CO; UT; Eurasia; Africa |
AK; AL; AR; AZ; CA; CO; CT; DE; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; QC; Worldwide
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Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In North America, as in Europe, Asplenium trichomanes consists of diploid and tetraploid cytotypes, treated here as subspecies. Asplenium trichomanes subsp. trichomanes, the diploid, is found on noncalcareous rocks. In the southwestern United States it occurs at high elevations. Asplenium trichomanes subsp. quadrivalens, the tetraploid, grows on calcareous substrates and has a more northern distribution (R. C. Moran 1982). Triploid hybrids are known between the diploids and tetraploids (R. C. Moran 1982; W. H. Wagner Jr. and F. S. Wagner 1966). Subspecies 4 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | |||||
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | Linnaeus: Sp. Pl. 2: 1080. (1753) | ||||
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