Asplenium adiantum-nigrum |
Asplenium resiliens |
|
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black spleenwort |
black-stem spleenwort |
|
Roots | not proliferous. |
not proliferous. |
Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
erect, unbranched; scales black throughout, linear-lanceolate, 4–5 × 0.2–0.6 mm, margins entire. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
black throughout, lustrous, 1.5–3(–5) cm, 1/4–1/10 length of blade; indument of blackish brown, filiform scales. |
Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
linear to narrowly oblanceolate, 1-pinnate throughout, 9–20(–30) × 1–2(–2.5) cm, thick, glabrous; base gradually tapered; apex acute, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
in 20–40 pairs, oblong; medial pinnae (5–)10–20 × (2–)3–5 mm; base usually with an acroscopic auricle; margins ± entire to shallowly crenate; apex obtuse. |
Veins | free, evident. |
free, obscure. |
Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
2–5 pairs per pinna, on both basiscopic and acroscopic sides, often confluent with age. |
Spores | 64 per sporangium. |
32 per sporangium. |
Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
black throughout, lustrous, glabrous. |
n | = 2n = 108 (apogamous). |
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2n | = 144. |
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Asplenium adiantum-nigrum |
Asplenium resiliens |
|
Habitat | Cliffs | Cliffs, sinkholes, on limestone or other basic rocks |
Elevation | 1675–2300 m (5500–7500 ft) | 100–1500 m (300–4900 ft) |
Distribution |
AZ; CO; UT; Eurasia; Africa |
AL; AR; AZ; DE; FL; GA; IL; KS; KY; LA; MD; MO; MS; NC; NM; NV; OK; PA; TN; TX; UT; VA; WV; Mexico; South America; West Indies in Hispaniola; Jamaica; Central America in Guatemala
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Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Asplenium parvulum M. Martens & Galeotti is an older, but illegitimate, name because it is a later homonym of A. parvulum Hooker. In Florida Asplenium resiliens hybridizes with A. heterochroum Kunze (4 x), producing A. × heteroresiliens (5 x). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | |
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | Kunze: Linnaea 18: 331. (1844) |
Web links |