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black spleenwort

black-stem spleenwort

Roots

not proliferous.

not proliferous.

Stems

ascending or short-creeping, infrequently branched;

scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate.

erect, unbranched;

scales black throughout, linear-lanceolate, 4–5 × 0.2–0.6 mm, margins entire.

Leaves

monomorphic.

monomorphic.

Petiole

dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade;

indument of black filiform scales and minute hairs.

black throughout, lustrous, 1.5–3(–5) cm, 1/4–1/10 length of blade;

indument of blackish brown, filiform scales.

Blade

deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute;

base truncate;

apex acute to acuminate, not rooting.

linear to narrowly oblanceolate, 1-pinnate throughout, 9–20(–30) × 1–2(–2.5) cm, thick, glabrous;

base gradually tapered;

apex acute, not rooting.

Pinnae

in 4–10 pairs, deltate to lanceolate;

most proximal (largest) pinnae 1.5–4 × 1–2.5 cm;

base obliquely obtuse;

segment margins coarsely incised;

apex acute.

in 20–40 pairs, oblong;

medial pinnae (5–)10–20 × (2–)3–5 mm;

base usually with an acroscopic auricle;

margins ± entire to shallowly crenate;

apex obtuse.

Veins

free, evident.

free, obscure.

Sori

1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides.

2–5 pairs per pinna, on both basiscopic and acroscopic sides, often confluent with age.

Spores

64 per sporangium.

32 per sporangium.

Rachis

greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent.

black throughout, lustrous, glabrous.

n

= 2n = 108 (apogamous).

2n

= 144.

Asplenium adiantum-nigrum

Asplenium resiliens

Habitat Cliffs Cliffs, sinkholes, on limestone or other basic rocks
Elevation 1675–2300 m (5500–7500 ft) 100–1500 m (300–4900 ft)
Distribution
from FNA
AZ; CO; UT; Eurasia; Africa
[BONAP county map]
from FNA
AL; AR; AZ; DE; FL; GA; IL; KS; KY; LA; MD; MO; MS; NC; NM; NV; OK; PA; TN; TX; UT; VA; WV; Mexico; South America; West Indies in Hispaniola; Jamaica; Central America in Guatemala
[WildflowerSearch map]
[BONAP county map]
Discussion

Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Asplenium parvulum M. Martens & Galeotti is an older, but illegitimate, name because it is a later homonym of A. parvulum Hooker.

In Florida Asplenium resiliens hybridizes with A. heterochroum Kunze (4 x), producing A. × heteroresiliens (5 x).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. andrewsii, A. chihuahuense, A. dubiosum
Name authority Linnaeus: Sp. Pl. 2: 1081. (1753) Kunze: Linnaea 18: 331. (1844)
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