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black spleenwort

bicolor spleenwort, varicolor spleenwort

Roots

not proliferous.

not proliferous.

Stems

ascending or short-creeping, infrequently branched;

scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate.

erect or ascending, rarely branched;

scales black throughout, linear-lanceolate, 2–3 × 0.2–0.4 mm, margin entire, apex attenuate.

Leaves

monomorphic.

monomorphic.

Petiole

dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade;

indument of black filiform scales and minute hairs.

black or purplish black throughout, lustrous, 0.3–5 cm, 1/4–1/15 length of blade;

indument of black filiform scales at base.

Blade

deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute;

base truncate;

apex acute to acuminate, not rooting.

linear or narrowly oblanceolate, 1-pinnate throughout, 6–22(–37) × 1–1.8(–2.3) cm, thin, glabrous;

base tapered;

apex acute, not rooting.

Pinnae

in 4–10 pairs, deltate to lanceolate;

most proximal (largest) pinnae 1.5–4 × 1–2.5 cm;

base obliquely obtuse;

segment margins coarsely incised;

apex acute.

in 15–40 pairs, oblong;

medial pinnae somewhat asymmetric, oblong, 4–10 × 2–4(–5) mm;

base acute, acroscopically enlarged;

margins serrate to crenate, lobed;

apex obtuse.

Veins

free, evident.

free, evident.

Sori

1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides.

3–6 pairs per pinna, on both basiscopic and acroscopic sides of pinnae.

Spores

64 per sporangium.

64 per sporangium.

Rachis

greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent.

black throughout, lustrous, glabrous.

2n

= 144.

= 144, 216.

Asplenium adiantum-nigrum

Asplenium heterochroum

Habitat Cliffs Sinkholes, limestone rocks in shady hammocks, masonry
Elevation 1675–2300 m (5500–7500 ft) 0–50 m (0–200 ft)
Distribution
from FNA
AZ; CO; UT; Eurasia; Africa
[BONAP county map]
from FNA
FL; GA; PR; Mexico; West Indies in Cuba; Bermuda; Central America in Belize
[BONAP county map]
Discussion

Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The tetraploid cytotype (4 x) of Asplenium heterochroum hybridizes with A. resiliens (3 x) to produce A. × heteroresiliens (5 x). It is extremely rare and local, known in northern peninsular Florida from Alachua, Citrus, Columbia, Jackson, and Marion counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. andrewsii, A. chihuahuense, A. dubiosum A. muticum
Name authority Linnaeus: Sp. Pl. 2: 1081. (1753) Kunze: Linnaea 9: 67. (1834)
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