Asplenium adiantum-nigrum |
Asplenium heterochroum |
|
---|---|---|
black spleenwort |
bicolor spleenwort, varicolor spleenwort |
|
Roots | not proliferous. |
not proliferous. |
Stems | ascending or short-creeping, infrequently branched; scales dark brown to blackish throughout, narrowly deltate, 2–4(–5) × 0.2–0.5 mm, margins entire or shallowly denticulate to serrulate. |
erect or ascending, rarely branched; scales black throughout, linear-lanceolate, 2–3 × 0.2–0.4 mm, margin entire, apex attenuate. |
Leaves | monomorphic. |
monomorphic. |
Petiole | dark reddish brown proximally, often fading to green distally, lustrous, 2–20 cm, 2/3–2 times length of blade; indument of black filiform scales and minute hairs. |
black or purplish black throughout, lustrous, 0.3–5 cm, 1/4–1/15 length of blade; indument of black filiform scales at base. |
Blade | deltate, 2–3-pinnate, 2.5–10 × 2–6.5 cm, thick, hairs dark, scattered, minute; base truncate; apex acute to acuminate, not rooting. |
linear or narrowly oblanceolate, 1-pinnate throughout, 6–22(–37) × 1–1.8(–2.3) cm, thin, glabrous; base tapered; apex acute, not rooting. |
Pinnae | in 4–10 pairs, deltate to lanceolate; most proximal (largest) pinnae 1.5–4 × 1–2.5 cm; base obliquely obtuse; segment margins coarsely incised; apex acute. |
in 15–40 pairs, oblong; medial pinnae somewhat asymmetric, oblong, 4–10 × 2–4(–5) mm; base acute, acroscopically enlarged; margins serrate to crenate, lobed; apex obtuse. |
Veins | free, evident. |
free, evident. |
Sori | 1–numerous pairs per pinna [1–6 pairs per segment], on both basiscopic and acroscopic sides. |
3–6 pairs per pinna, on both basiscopic and acroscopic sides of pinnae. |
Spores | 64 per sporangium. |
64 per sporangium. |
Rachis | greenish throughout or sometimes reddish brown proximally, lustrous, sparsely pubescent. |
black throughout, lustrous, glabrous. |
2n | = 144. |
= 144, 216. |
Asplenium adiantum-nigrum |
Asplenium heterochroum |
|
Habitat | Cliffs | Sinkholes, limestone rocks in shady hammocks, masonry |
Elevation | 1675–2300 m (5500–7500 ft) | 0–50 m (0–200 ft) |
Distribution |
AZ; CO; UT; Eurasia; Africa |
FL; GA; PR; Mexico; West Indies in Cuba; Bermuda; Central America in Belize |
Discussion | Asplenium adiantum-nigrum is principally a Eurasian species and occurs extremely rarely in North America (see M. G. Shivas 1969 and M. D. Windham 1983 for a discussion of the conspecificity of Western Hemisphere and Eastern Hemisphere material). It is an allotetraploid derived from hybridization of two European taxa, A. cuneifolium Viviani and A. onopteris Linnaeus (M. G. Shivas 1969). Hybrids involving A. adiantum-nigrum and other Asplenium species occur in Europe but are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The tetraploid cytotype (4 x) of Asplenium heterochroum hybridizes with A. resiliens (3 x) to produce A. × heteroresiliens (5 x). It is extremely rare and local, known in northern peninsular Florida from Alachua, Citrus, Columbia, Jackson, and Marion counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Aspleniaceae > Asplenium | Aspleniaceae > Asplenium |
Sibling taxa | ||
Synonyms | A. andrewsii, A. chihuahuense, A. dubiosum | A. muticum |
Name authority | Linnaeus: Sp. Pl. 2: 1081. (1753) | Kunze: Linnaea 9: 67. (1834) |
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