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aspidote touffue, dense lace fern, Indian's-dream, lace fern, Oregon cliff brake, podfern

aspidote, aspidotis, Indian's dream, lace ferns, lacefern

Habit Plants terrestrial or on rock.
Stems

± compact, short-creeping, ascending at tip, branched;

scales mostly dark brown, often with very narrow margin of lighter color, lanceolate, margins entire.

Leaves

monomorphic or often somewhat dimorphic, 8–25 cm;

fertile leaves more erect than sterile leaves, long-petioled, petioles often 2–5 times longer than blades, fertile blades with more ascending pinnae and narrower segments than sterile blades.

monomorphic to somewhat dimorphic, crowded, 8–35 cm.

Petiole

usually dark reddish brown, with single groove adaxially, glabrous, with single vascular bundle.

Blade

3–4-pinnate, 2–10 cm, somewhat leathery.

ovate-triangular, deltate, or pentagonal, 3–4(–5)-pinnate, thick to thin, abaxially glabrous, adaxially lustrous, striate, glabrous;

rachis straight.

Ultimate segments

linear, 3–8 mm;

midrib prominent abaxially.

of blades short-stalked or with base narrowed and decurrent onto costa or costule-bearing segments, linear to lanceolate, mostly 0.5–1.3 mm wide;

stalks greenish, not darkened;

fertile margins recurved.

Veins

of ultimate segments obscure, free, ± pinnate and unbranched.

False indusia

appearing inframarginal, scarious, whitish, broad, partly concealing sporangia.

Sori

of mature blades continuous along length of segments except at apex;

indusia linear, margins with 10–35, shallow, regular teeth or erose.

Sporangia

in marginal, discrete or continuous sori on abaxial surface, containing 64 spores, lacking paraphyses and glands.

Spores

dark brown, tetrahedral-globose, trilete, reticulate, equatorial flange absent.

x

= 30.

2n

= 60.

Aspidotis densa

Aspidotis

Habitat Slopes, crevices, rocky outcrops, often on serpentine, sometimes in chaparral
Elevation 300–3400 m (1000–11200 ft)
Distribution
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; BC; QC
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; 1 in Mexico
[BONAP county map]
Discussion

D. B. Lellinger (1968) recognized Aspidotis as separate from Cheilanthes based on its elongate, distantly dentate segments with striate shining surface and on its broad, scarious indusia.

Species 4 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Mature, fertile blades with continuous sori along length of segments (not at apex); indusia with 10–35 shallow, regular teeth or erose; fertile segments linear, margins ± entire.
A. densa
1. Mature, fertile blades with sori discrete or partially discontinuous; indusia with coarse, irregular teeth or entire; fertile segments lanceolate to deltate, distantly dentate.
→ 2
2. Sori discrete, 1–3(–5) per blade segment; indusia margins with 2–6 coarse, irregular teeth or ± entire.
A. californica
2. Sori partially discontinuous, connected by narrow indusial wings, 3–7(–9) per blade segment; indusia margins with 6–10 coarse, irregular teeth or lobes.
A. carlotta-halliae
Source FNA vol. 2. FNA vol. 2. Author: Alan R. Smith.
Parent taxa Pteridaceae > Aspidotis Pteridaceae
Sibling taxa
A. californica, A. carlotta-halliae
Subordinate taxa
A. californica, A. carlotta-halliae, A. densa
Synonyms Onychium densum, Cheilanthes siliquosa, Pellaea densa Hypolepis section A.
Name authority (Brackenridge in Wilkes) Lellinger: Amer. Fern J. 58: 141. (1968) (Nuttall ex Hooker & Baker) Copeland: Gen. Fil. 68. (1947)
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