Asclepias cordifolia |
Asclepias albicans |
|
---|---|---|
heart-leaf milkweed, purple milkweed |
candelilla, wax milkweed, white-stem milkweed |
|
Habit | Herbs. | Shrubs. |
Stems | 1–20, ascending to spreading, unbranched, 25–100 cm, glabrous, glaucous, rhizomes absent. |
5–40 (usually 1–few), erect to ascending, branched, especially in lower half, 140–400 cm, sparsely pilose to glabrate, thickly glaucous, rhizomes absent. |
Leaves | opposite, sessile, stipular colleters absent; blade ovate to lanceolate, 3.5–11 × 1.8–8 cm, chartaceous, base cordate, clasping, margins entire, apex acute, mucronate, venation brochidodromous to eucamptodromous, surfaces glabrous, glaucous, margins minutely ciliate, laminar colleters absent. |
ephemeral, rarely present on flowering stems, opposite, sessile, with 0 or 1 stipular colleter on each side of leaf base; blade filiform, 1.5–2.5 × 0.1 cm, succulent, base cuneate, margins entire, apex acute, mucronate, venation obscure, surfaces pilosulous, laminar colleters absent. |
Inflorescences | terminal, branched, sometimes also extra-axillary at upper nodes, pedunculate, 5–20-flowered; peduncle 0.3–6.5 cm, apically sparsely pilose to glabrate, with few bracts. |
terminal, branched, also extra-axillary at leafless upper nodes, pedunculate, 8–50-flowered; peduncle 0.2–4.5 cm, pilose, with 1 caducous bract at the base of each pedicel. |
Pedicels | 16–37 mm, sparsely pilose. |
8–16 mm, pilose. |
Flowers | erect to pendent; calyx lobes elliptic to lanceolate, 3–5 mm, apex acute, pilose; corolla red-violet, lobes reflexed with spreading tips, oval, 6–7 mm, apex acute, glabrous; gynostegial column 1–1.5 mm; fused anthers brown, cylindric, 1.5–2 mm, wings right-triangular, open at tip, apical appendages deltoid; corona segments white, tinged pink to red-violet at base, sessile, tubular, 2–3 mm, exceeded by style apex, base saccate, apex truncate, oblique, glabrous, internal appendage absent; style apex shallowly depressed, pink to red-violet. |
erect to pendent; calyx lobes lanceolate to linear-lanceolate, 2.5–3 mm, apex acute, densely pilose; corolla ochroleucous to greenish cream, sometimes tinged red, faintly striate, lobes reflexed or sometimes spreading, oval, 4.5–6 mm, apex acute, glabrous; gynostegial column 1–1.8 mm; fused anthers brown, cylindric, 1.8–2 mm, wings right-triangular, closed, apical appendages ovate, erose; corona segments cream, often tinged pink, yellow, or green, shiny, sessile, conduplicate, 2–3 mm, exceeded by style apex, base saccate, apex truncate, oblique, glabrous, internal appendage falcate, exserted, sharply inflexed towards style apex, densely papillate; style apex shallowly depressed, ochroleucous to green. |
Seeds | ovate, 8–9 × 5–7 mm, margin not winged, faces reticulate-rugulose; coma 3–3.5 cm. |
naviculate, lanceolate, 5–7 × 2.5–3 mm, margin narrowly winged, faces papillose and rugulose, concave face with a low keel; coma 1.5–2 cm. |
Follicles | erect on upcurved pedicels, lance-ovoid, 7.5–10.5 × 1.5–1.8 cm, apex long-acuminate, smooth, glabrous, glaucous. |
spreading to pendulous on spreading to pendulous pedicels, fusiform to lance-ovoid, 5.5–12 × 0.7–1.8 cm, apex acuminate, smooth, pilosulous. |
2n | = 22. |
|
Asclepias cordifolia |
Asclepias albicans |
|
Phenology | Flowering Mar–Jul(–Aug); fruiting Jun–Sep. | Flowering and fruiting year-round. |
Habitat | Hillsides, canyons, ridge tops, streamsides, seeps, basalt, serpentine, gabbro, granite, shale, limestone, talus slopes, gravel, alluvium, oak woodlands, mixed evergreen, douglas-fir, pine, pine-oak, and riparian forests, chaparral, timberline meadows, grasslands. | Mountain slopes, ridge tops, bajadas, flats, arroyos, granite, basalt, tuff, coarse rocky soils, cracks in boulders, sand, desert scrub. |
Elevation | 50–2200(–2800) m. [160–7200(–9200) ft.] | 50–800 m. [160–2600 ft.] |
Distribution |
CA; NV; OR
|
AZ; CA; Mexico (Baja California, Baja California Sur, Sonora)
|
Discussion | Fresh leaves and stems of Asclepias cordifolia are often slightly or strongly colored bluish, grayish, or purplish. This is one of the few American species of Asclepias with cavitate corona segments that lack adaxial appendages. Such species were segregated along with diverse African species in Gomphocarpus R. Brown, a polyphyletic segregate (M. Fishbein et al. 2011; D. Chuba et al. 2017). Asclepias cordifolia is a distinctive species unlike any other within its range. It is phylogenetically and geographically isolated, although not highly derived morphologically. In Nevada, A. cordifolia is restricted to the Sierra Nevada, in Carson City, Douglas, and Washoe counties. Its range extends to northern California and southwestern Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Asclepias albicans reaches the greatest height of any Asclepias species in the flora area and among congeners can be confused only with A. subulata, the only other shrubby, leafless milkweed in the region. In the absence of flowers, it may be difficult to distinguish these species. In addition to the key characters, A. albicans often differs by thicker, waxier stems. The species are also ecologically divergent: A. albicans is most commonly found on rocky slopes and ridges (usually basalt or granite), and A. subulata is usually found on flats and in arroyos, often in sandy soils. However, A. albicans can occur on bajadas and in arroyos, often distant from the mountains and ridges harboring source populations. These species have extensively overlapping ranges but rarely hybridize at widely scattered locations in Arizona, California, and Baja California Sur. Hybrids are identified by intermediate floral morphology, especially corona size and shape. The range of A. albicans is within the limits of the Sonoran Desert. In Arizona, it is found in La Paz, Maricopa, Mohave, Pima, Pinal, and Yuma counties; in California only in Imperial, Riverside, San Bernardino, and San Diego counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 14. | FNA vol. 14. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Acerates cordifolia, Gomphocarpus cordifolius | |
Name authority | (Bentham) Jepson: Fl. W. Calif., 384. (1901) | S. Watson: Proc. Amer. Acad. Arts 24: 59. (1889) |
Web links |