Asclepias cordifolia |
Asclepias variegata |
|
---|---|---|
heart-leaf milkweed, purple milkweed |
asclépiade panachée, red-ringed milkweed, redring milkweed, white milkweed |
|
Habit | Herbs. | Herbs. |
Stems | 1–20, ascending to spreading, unbranched, 25–100 cm, glabrous, glaucous, rhizomes absent. |
1–few, erect, unbranched, 30–120 cm, puberulent with curved trichomes, sometimes in a single line, sometimes glaucous, rhizomes absent. |
Leaves | opposite, sessile, stipular colleters absent; blade ovate to lanceolate, 3.5–11 × 1.8–8 cm, chartaceous, base cordate, clasping, margins entire, apex acute, mucronate, venation brochidodromous to eucamptodromous, surfaces glabrous, glaucous, margins minutely ciliate, laminar colleters absent. |
opposite, petiolate, with 1 or 2 stipular colleters on each side of petiole; petiole 10–25 mm, puberulent with curved trichomes; blade oval to ovate, obovate, lanceolate, or oblanceolate, 6–15 × 3–9 cm, subcoriaceous, base cuneate to obtuse or subtruncate, margins sometimes crisped, apex rounded to obtuse, apiculate or mucronate, venation eucamptodromous to faintly brochidodromous, surfaces puberulent on veins with curved trichomes, sometimes glaucous, margins ciliate, 8–12 laminar colleters. |
Inflorescences | terminal, branched, sometimes also extra-axillary at upper nodes, pedunculate, 5–20-flowered; peduncle 0.3–6.5 cm, apically sparsely pilose to glabrate, with few bracts. |
terminal, branched, also usually extra-axillary at 1 distal node, pedunculate, 11–28-flowered; peduncle 1–7 cm, densely puberulent with curved trichomes, with 1 caducous bract at the base of each pedicel. |
Pedicels | 16–37 mm, sparsely pilose. |
12–20 mm, densely puberulent with curved trichomes. |
Flowers | erect to pendent; calyx lobes elliptic to lanceolate, 3–5 mm, apex acute, pilose; corolla red-violet, lobes reflexed with spreading tips, oval, 6–7 mm, apex acute, glabrous; gynostegial column 1–1.5 mm; fused anthers brown, cylindric, 1.5–2 mm, wings right-triangular, open at tip, apical appendages deltoid; corona segments white, tinged pink to red-violet at base, sessile, tubular, 2–3 mm, exceeded by style apex, base saccate, apex truncate, oblique, glabrous, internal appendage absent; style apex shallowly depressed, pink to red-violet. |
erect to spreading; calyx lobes lanceolate, 2–3 mm, apex acute, pilosulous; corolla white, red-violet in throat, lobes reflexed with spreading tips, elliptic, 6–8 mm, apex acute, glabrous abaxially, minutely papillose at base adaxially; gynostegial column red-violet, 1–2 mm; fused anthers brown, truncately obconic, 1.5–2 mm, wings right-triangular, rounded, closed, apical appendages ovate; corona segments white, stipitate, conduplicate with a lateral flange on each side, 2.5–4 mm, exceeding style apex, apex truncate, glabrous, internal appendage falcate, exserted, sharply inflexed over style apex, glabrous; style apex shallowly depressed, white. |
Seeds | ovate, 8–9 × 5–7 mm, margin not winged, faces reticulate-rugulose; coma 3–3.5 cm. |
ovate, 5–7 × 3–5 mm, margin winged, faces rugulose; coma 2.5–4 cm. |
Follicles | erect on upcurved pedicels, lance-ovoid, 7.5–10.5 × 1.5–1.8 cm, apex long-acuminate, smooth, glabrous, glaucous. |
erect on upcurved pedicels, narrowly fusiform, 10–15 × 1.5–2 cm, apex long-acuminate, smooth, pilosulous. |
2n | = 22. |
|
Asclepias cordifolia |
Asclepias variegata |
|
Phenology | Flowering Mar–Jul(–Aug); fruiting Jun–Sep. | Flowering Mar–Aug; fruiting Jun–Nov. |
Habitat | Hillsides, canyons, ridge tops, streamsides, seeps, basalt, serpentine, gabbro, granite, shale, limestone, talus slopes, gravel, alluvium, oak woodlands, mixed evergreen, douglas-fir, pine, pine-oak, and riparian forests, chaparral, timberline meadows, grasslands. | Ridges, slopes, bluffs, flats, glades, ravines, streamsides, lake shores, limestone, sandstone, basalt, clay, sandy, silty, and marl soils, oak-hickory, oak, mixed-hardwood, pine-mixed-hardwood, pine-oak-hickory, and pine forests, oak, pine, pine-oak, oak-hickory, and riparian woodlands, forest edges and openings. |
Elevation | 50–2200(–2800) m. (200–7200(–9200) ft.) | 0–900 m. (0–3000 ft.) |
Distribution |
CA; NV; OR
|
AL; AR; CT; DC; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; ON
|
Discussion | Fresh leaves and stems of Asclepias cordifolia are often slightly or strongly colored bluish, grayish, or purplish. This is one of the few American species of Asclepias with cavitate corona segments that lack adaxial appendages. Such species were segregated along with diverse African species in Gomphocarpus R. Brown, a polyphyletic segregate (M. Fishbein et al. 2011; D. Chuba et al. 2017). Asclepias cordifolia is a distinctive species unlike any other within its range. It is phylogenetically and geographically isolated, although not highly derived morphologically. In Nevada, A. cordifolia is restricted to the Sierra Nevada, in Carson City, Douglas, and Washoe counties. Its range extends to northern California and southwestern Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Asclepias variegata and the following species (A. exaltata, A. purpurascens, and A. quadrifolia) inhabit deciduous forest understories in eastern North America and often co-occur. Asclepias variegata has showy, snowball-like spheres of bright white flowers. On closer examination, the staminal column of each flower is colored reddish purple, forming a neat belt between the corona and corolla. Non-flowering specimens have been confused with A. purpurascens. The leaves of A. variegata have a thicker texture, and the blade apices are broader and more rounded than in A. purpurascens. Asclepias variegata has suffered serious declines at the northeastern margin of its range and is reported to have been extirpated from Ontario and Connecticut. In addition, it is considered to be of conservation concern in Delaware, New Jersey, New York, and Pennsylvania. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 14. | FNA vol. 14. |
Parent taxa | Apocynaceae > Asclepias | Apocynaceae > Asclepias |
Sibling taxa | ||
Synonyms | Acerates cordifolia, Gomphocarpus cordifolius | Biventraria variegata |
Name authority | (Bentham) Jepson: Fl. W. Calif., 384. (1901) | Linnaeus: Sp. Pl. 1: 215. (1753) |
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