Asclepias cordifolia |
Asclepias asperula |
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heart-leaf milkweed, purple milkweed |
Antelope horns, spider milkweed |
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Habit | Herbs. | Herbs. | ||||
Stems | 1–20, ascending to spreading, unbranched, 25–100 cm, glabrous, glaucous, rhizomes absent. |
1–40, erect to decumbent, unbranched or branched at base, 15–60 cm, puberulent with curved trichomes, not glaucous, rhizomes absent. |
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Leaves | opposite, sessile, stipular colleters absent; blade ovate to lanceolate, 3.5–11 × 1.8–8 cm, chartaceous, base cordate, clasping, margins entire, apex acute, mucronate, venation brochidodromous to eucamptodromous, surfaces glabrous, glaucous, margins minutely ciliate, laminar colleters absent. |
alternate to subopposite, petiolate, with 1–3 stipular colleters on each side of petiole; petiole 2–4 mm, puberulent with curved trichomes to pilosulous; blade lanceolate to linear, 5–17 × 0.4–3.7 cm, chartaceous, base cuneate, margins entire, apex attenuate to acute, mucronate, venation eucamptodromous to brochidodromous, surfaces puberulent with curved trichomes to glabrate, more densely so on veins, margins ciliate, laminar colleters absent. |
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Inflorescences | terminal, branched, sometimes also extra-axillary at upper nodes, pedunculate, 5–20-flowered; peduncle 0.3–6.5 cm, apically sparsely pilose to glabrate, with few bracts. |
terminal, sessile or pedunculate, 10–60-flowered; peduncle 0–22.5 cm, puberulent with curved trichomes, with 1 caducous bract at the base of each pedicel. |
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Pedicels | 16–37 mm, sparsely pilose. |
16–30 mm, puberulent with curved trichomes to pilose. |
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Flowers | erect to pendent; calyx lobes elliptic to lanceolate, 3–5 mm, apex acute, pilose; corolla red-violet, lobes reflexed with spreading tips, oval, 6–7 mm, apex acute, glabrous; gynostegial column 1–1.5 mm; fused anthers brown, cylindric, 1.5–2 mm, wings right-triangular, open at tip, apical appendages deltoid; corona segments white, tinged pink to red-violet at base, sessile, tubular, 2–3 mm, exceeded by style apex, base saccate, apex truncate, oblique, glabrous, internal appendage absent; style apex shallowly depressed, pink to red-violet. |
erect to spreading; calyx lobes ovate to linear-lanceolate, 3–5 mm, apex acute, pilosulous to puberulent with curved trichomes; corolla pale green, sometimes tinged red abaxially, campanulate, lobes ascending and exceeding corolla segments, ovate to oval, 7–10 mm, apex acute, puberulent with curved trichomes at apex abaxially, glabrous adaxially; gynostegium sessile; fused anthers brown and green, turbinate, 2–2.5 mm, wings trapezoidal, widest above middle, closed, apical appendages ovate, erose; corona segments reddish purple and white, sessile, clavate-tubular, 4.5–7 mm, slightly exceeded by to equaling style apex, deflexed at base, margins connivent, apex incurved, rounded, upper margin and cavity papillose, internal appendage a low internal crest, papillose; style apex depressed, green. |
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Seeds | ovate, 8–9 × 5–7 mm, margin not winged, faces reticulate-rugulose; coma 3–3.5 cm. |
ovate, 5–8 × 4–6 mm, margin winged, remotely erose, faces minutely rugulose-papillose, minutely hirtellous; coma 2.5–4 cm. |
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Follicles | erect on upcurved pedicels, lance-ovoid, 7.5–10.5 × 1.5–1.8 cm, apex long-acuminate, smooth, glabrous, glaucous. |
erect on upcurved pedicels, lance-ovoid, 6–11.5 × 1–2.5 cm, apex short- to long-acuminate, weakly to strongly arcuate, shallowly rugose-ribbed, ribs sometimes muricate, striate, pilosulous. |
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2n | = 22. |
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Asclepias cordifolia |
Asclepias asperula |
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Phenology | Flowering Mar–Jul(–Aug); fruiting Jun–Sep. | |||||
Habitat | Hillsides, canyons, ridge tops, streamsides, seeps, basalt, serpentine, gabbro, granite, shale, limestone, talus slopes, gravel, alluvium, oak woodlands, mixed evergreen, douglas-fir, pine, pine-oak, and riparian forests, chaparral, timberline meadows, grasslands. | |||||
Elevation | 50–2200(–2800) m. (200–7200(–9200) ft.) | |||||
Distribution |
CA; NV; OR
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w United States; Mexico
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Discussion | Fresh leaves and stems of Asclepias cordifolia are often slightly or strongly colored bluish, grayish, or purplish. This is one of the few American species of Asclepias with cavitate corona segments that lack adaxial appendages. Such species were segregated along with diverse African species in Gomphocarpus R. Brown, a polyphyletic segregate (M. Fishbein et al. 2011; D. Chuba et al. 2017). Asclepias cordifolia is a distinctive species unlike any other within its range. It is phylogenetically and geographically isolated, although not highly derived morphologically. In Nevada, A. cordifolia is restricted to the Sierra Nevada, in Carson City, Douglas, and Washoe counties. Its range extends to northern California and southwestern Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). With terminal umbels of large, green and purple flowers, Asclepias asperula is highly distinctive and in flower can be confused only with its sister species, A. viridis. Distinguishing characteristics and the existence of interspecific hybrids are discussed under A. viridis. Large bees, notably Bombus and Xylocopa, are commonly observed visiting the flowers of A. asperula. The subspecies of Asclepias asperula are strongly differentiated away from their region of contact, which extends from south of the Texas Panhandle to the extreme tip of the Oklahoma Panhandle. In the region of contact, their distinguishing traits intermix. The common occurrence of intermediates and apparent introgressants argues against elevation of the subspecies to the specific rank. There is a surprising gap in the distribution of the species as a whole on the Llano Estacado in eastern New Mexico and the Texas Panhandle that may contribute to the differentiation of the subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 14. | FNA vol. 14. | ||||
Parent taxa | Apocynaceae > Asclepias | Apocynaceae > Asclepias | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Acerates cordifolia, Gomphocarpus cordifolius | Acerates asperula, Asclepiodora asperula | ||||
Name authority | (Bentham) Jepson: Fl. W. Calif., 384. (1901) | (Decaisne) Woodson: Ann. Missouri Bot. Gard. 41: 193. (1954) | ||||
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