Asclepias albicans
Asclepias arenaria
candelilla, wax milkweed, white-stem milkweed
sand milkweed
5–40 (usually 1–few), erect to ascending, branched, especially in lower half, 140–400 cm, sparsely pilose to glabrate, thickly glaucous, rhizomes absent.
1–8, spreading or decumbent to erect, unbranched or rarely branched near base, 20–100 cm, tomentose, not glaucous, rhizomatous.
ephemeral, rarely present on flowering stems, opposite, sessile, with 0 or 1 stipular colleter on each side of leaf base;
blade filiform, 1.5–2.5 × 0.1 cm, succulent, base cuneate, margins entire, apex acute, mucronate, venation obscure, surfaces pilosulous, laminar colleters absent.
opposite, petiolate, with 1 or 2 stipular colleters on each side of petiole plus 0–4 in axil;
petiole 7–17 mm, tomentose;
blade oblong or obovate to ovate or oval, 4.2–11.5 × 2.5–7.5 cm, subcoriaceous, base rounded or truncate to subcordate, margins often undulate or crisped, apex truncate to rounded (rarely acute), sometimes emarginate, often mucronate, venation brochidodromous, surfaces tomentose to nearly glabrate, margins ciliate, 12–24 laminar colleters.
terminal, branched, also extra-axillary at leafless upper nodes, pedunculate, 8–50-flowered;
peduncle 0.2–4.5 cm, pilose, with 1 caducous bract at the base of each pedicel.
extra-axillary (sometimes appearing terminal), sessile or pedunculate, 14–51-flowered;
peduncle 0–2 cm, tomentose, with 1 caducous bract at the base of each pedicel.
8–16 mm, pilose.
15–25 mm, densely tomentose.
erect to pendent;
calyx lobes lanceolate to linear-lanceolate, 2.5–3 mm, apex acute, densely pilose;
corolla ochroleucous to greenish cream, sometimes tinged red, faintly striate, lobes reflexed or sometimes spreading, oval, 4.5–6 mm, apex acute, glabrous;
gynostegial column 1–1.8 mm;
fused anthers brown, cylindric, 1.8–2 mm, wings right-triangular, closed, apical appendages ovate, erose;
corona segments cream, often tinged pink, yellow, or green, shiny, sessile, conduplicate, 2–3 mm, exceeded by style apex, base saccate, apex truncate, oblique, glabrous, internal appendage falcate, exserted, sharply inflexed towards style apex, densely papillate;
style apex shallowly depressed, ochroleucous to green.
erect to pendent;
calyx lobes lanceolate, 2.5–3 mm, apex acute, densely tomentose;
corolla green to yellowish green, sometimes tinged reddish or purplish, lobes reflexed with spreading tips, oval, 7–8 mm, apex acute, glabrous abaxially, minutely papillose at base adaxially;
gynostegial column 1–2 mm;
fused anthers green, obconic, 2–2.5 mm, wings right-triangular, closed, apical appendages ovate;
corona segments cream to greenish cream or ochroleucous, subsessile, conduplicate, flaring at base, 3.5–4 mm, exceeding style apex, apex truncate to rounded, emarginate, minutely papillate, proximal margin toothed, internal appendage falcate, exserted, sharply incurved over style apex, apex upturned, minutely papillose.
naviculate, lanceolate, 5–7 × 2.5–3 mm, margin narrowly winged, faces papillose and rugulose, concave face with a low keel;
coma 1.5–2 cm.
oval, 9–12 × 6–8 mm, margin winged, faces minutely rugulose;
coma 2–3 cm.
spreading to pendulous on spreading to pendulous pedicels, fusiform to lance-ovoid, 5.5–12 × 0.7–1.8 cm, apex acuminate, smooth, pilosulous.
erect on upcurved pedicels, lance-ovoid, 5.5–10 × 2–2.8 cm, apex acuminate, smooth, pilosulous.
Asclepias albicans
Asclepias arenaria
Asclepias albicans reaches the greatest height of any Asclepias species in the flora area and among congeners can be confused only with A. subulata, the only other shrubby, leafless milkweed in the region. In the absence of flowers, it may be difficult to distinguish these species. In addition to the key characters, A. albicans often differs by thicker, waxier stems. The species are also ecologically divergent: A. albicans is most commonly found on rocky slopes and ridges (usually basalt or granite), and A. subulata is usually found on flats and in arroyos, often in sandy soils. However, A. albicans can occur on bajadas and in arroyos, often distant from the mountains and ridges harboring source populations. These species have extensively overlapping ranges but rarely hybridize at widely scattered locations in Arizona, California, and Baja California Sur. Hybrids are identified by intermediate floral morphology, especially corona size and shape. The range of A. albicans is within the limits of the Sonoran Desert. In Arizona, it is found in La Paz, Maricopa, Mohave, Pima, Pinal, and Yuma counties; in California only in Imperial, Riverside, San Bernardino, and San Diego counties.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Asclepias arenaria is the milkweed most consistently associated with pure sand soils in the western Great Plains. It is predictably found at the bases of stabilized and semi-stabilized dunes. Flowers of this species are visited by a variety of Hymenoptera, notably several species of large wasps, including tarantula hawk wasps (Pepsinae, Pompilidae) and scoliid wasps (Scoliidae), as well as by Lepidoptera. Non-flowering shoots of A. arenaria may produce linear leaves; they are easily overlooked and not identified as belonging to this species unless one is aware of this variation, especially when they are produced on rhizomes distant from shoots with typical foliage. This trait is found in several other broad-leaved milkweeds (for example, A. erosa, A. welshii). Asclepias arenaria is rare and considered to be of conservation concern in Wyoming, where it has been recorded from only two sites in Goshen County.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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