armoise de Steller, beach wormwood, dusty miller, oldwoman, Steller's wormwood
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Perennials, (15–)20–60(–70) cm (mat-forming), sometimes faintly aromatic (rhizomes creeping, relatively thin). |
Annuals, biennials, or perennials (shrubs in A. californica, subshrubs in A. nesiotica); usually fibrous-rooted, sometimes taprooted, caudices sometimes woody, rhizomes sometimes present. |
1–3, erect or ascending, white, simple (stout), densely tomentose to floccose. |
usually not wandlike (wandlike in A. californica, A. nesiotica, A. palmeri). |
basal and cauline (petiolate), silver-gray; blades oblanceolate, (proximalmost) 3–10 × 1–5 cm, pinnatifid (lobes relatively broad, rounded; distal leaves, on flowering stems, smaller), faces densely tomentose. |
usually deciduous, rarely persistent, basal (rosettes) and/or cauline (not in fascicles). |
broadly campanulate, 5–8 × 6–7 mm. |
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glabrous (paleate in A. palmeri). |
pistillate 12–16; bisexual 25–30; corollas yellow (narrow or tubular), 3.2–4 mm (unusually large), glabrous or sparsely hairy (style branches prominent, erect, blunt). |
usually peripheral 3–20 pistillate and fertile (0 pistillate in A. nesiotica, A. palmeri); central (or all) 14–70 bisexual and fertile; corollas funnelform. |
broadly lanceolate, tomentose. |
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(erect or spreading, peduncles 0 or to 3 mm) in dense, paniculiform, racemiform, or spiciform arrays 8–20 × 2–4 cm. |
usually disciform (discoid in A. nesiotica and A. palmeri). |
(dark brown) narrowly oblong-linear (slightly flattened, smooth), 3–4 mm, glabrous. |
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= 18. |
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Flowering early spring–fall. |
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Sandy soils, coastal strand |
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0–200 m (0–700 ft) |
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AK; CT; DE; FL; LA; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; VA; VT; WA; WI; WV; NB; NF; NS; ON; PE; QC; SPM; n Europe; e Asia (Japan, Kamchatka)
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Widespread in Northern Hemisphere; especially North America; Europe; and central and northern Asia; sporadic in South America and northern Africa |
Artemisia stelleriana is apparently native along the western tip of the Aleutian islands (D. F. Murray, pers. comm.). It is an attractive ornamental and, in parts of its range in the flora area, it appears to have escaped from cultivation and is naturalized in beach dunes and other sandy habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 220 (27 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Subshrubs or shrubs (stems wandlike) | → 2 |
1. Annuals, biennials, perennials, subshrubs, or shrubs (stems sometimes brittle, not wandlike) | → 4 |
2. Plants 100–350 cm; leaves relatively deeply and coarsely pinnately lobed (lobes 3–7+; coastal California and Baja California) | A. palmeri |
2. Plants 10–250 cm; leaves pinnately lobed or 3-lobed | → 3 |
3. Shrubs (20–250 cm); leaves pinnately lobed (lobes 0.5–1 mm wide); California (chaparral) | A. californica |
3. Subshrubs (10–60 cm, stems mostly prostrate); leaves 3-lobed (lobes 1–2 mm wide); Channel Islands, California | A. nesiotica |
4. Annuals or biennials; leaves among heads (relatively deeply) lobed | → 5 |
4. Perennials, subshrubs, or shrubs; leaves among heads mostly entire | → 6 |
5. Annuals, 30–200(–300) cm, sweetly aromatic; leaves 2–3-pinnatifid; arrays of heads 10–20 cm diam | A. annua |
5. Annuals or biennials, (10–)30–80(–150) cm, not aromatic; leaves 1–2-pinnately lobed; arrays of heads 2–4 cm diam | A. biennis |
6. Perennials, subshrubs, or shrubs (not rhizomatous) | → 7 |
6. Perennials (usually rhizomatous, stems sometimes woody at bases) | → 15 |
7. Heads in capitate or dense, corymbiform arrays (plants cespitose) | → 8 |
7. Heads in paniculiform or racemiform arrays (plants not cespitose) | → 10 |
8. Leaves pinnately lobed; involucres 3–5 mm diam.; phyllaries lanceolate to ovate (margins white) | A. senjavinensis |
8. Leaves 1–2-palmatifid; involucres 3.5–11 mm diam.; phyllaries lanceolate (margins brown or white) | → 9 |
9. Involucres 3.5–6 × 6–11 mm; phyllaries lanceolate (margins brown); corollas yellow or reddish black, glabrous or glandular (not pilose). | A. globularia |
9. Involucres 3–4 × 3.5–5 mm; phyllaries lanceolate (margins white); corollas yellow, glabrous or pilose | A. glomerata |
10. Leaves entire, irregularly palmatifid, or palmately 3-lobed to 2-ternately lobed | → 11 |
10. Leaves 2–3-pinnatifid | → 13 |
11. Pappi coroniform; Idaho, Nevada, Oregon | A. papposa |
11. Pappi 0; Alberta, British Columbia, Northwest Territories, Nunavut; Alaska, Washington | → 12 |
12. Plants 15–70 cm; leaves palmately 3-lobed to 2-ternately lobed. | A. alaskana |
12. Plants (5–)10–40 cm; leaves 1–3-palmately lobed | A. furcata |
13. Perennials or subshrubs, 50–170 cm (widely branched, stems brittle); leaf lobes less than 1 mm wide; heads erect; involucres 1.5–3 mm diam. (gardens, waste places, much of North America) | A. abrotanum |
13. Perennials, 10–50 cm (erect, stems not brittle); leaf lobes 1+ mm wide (margins coarsely toothed); heads nodding; involucres 4–10 mm diam | → 14 |
14. Peduncles 0 or to 10 mm | A. laciniata |
| A. norvegica |
15. Leaves entire, serrate, toothed, or lobed (sinuses to 1/2 blade widths) | → 16 |
15. Leaves (relatively deeply) lobed (sinuses 1/2+ blade widths) | → 22 |
16. Plants 20–50(–80, rarely more) cm | → 17 |
| → 19 |
17. Leaves usually entire, sometimes toothed or lobed; involucres 4–5 mm diam | A. longifolia |
17. Leaves usually lobed, sometimes entire; involucres (1–)2–5 mm diam | → 18 |
18. Involucres 2–4(–5.5) mm; phyllaries (gray-green) densely tomentose | A. ludoviciana |
18. Involucres 3.5–4 mm; phyllaries (violet-brown) sparsely tomentose (w North America) | A. tilesii |
19. Phyllaries glabrous or sparsely hairy (coast, n California to British Columbia) | A. suksdorfii |
19. Phyllaries usually densely hairy | → 20 |
20. Leaves densely hairy (both faces, broadly lanceolate, mostly entire, the proximal lobed; w North America, mostly inland grasslands). | A. douglasiana |
20. Leaves (bicolor) hairy abaxially, glabrate or glabrous adaxially | → 21 |
21. Leaves serrate (teeth ca. 2 mm; inland grasslands and barren areas, high plains) | A. serrata |
21. Leaves mostly deeply lobed (lobes 4–20 mm; mostly e North America, introduced w coast) | A. vulgaris |
22. Leaves not bicolor (both faces bright green or silvery), lobes acute or rounded | → 23 |
22. Leaves bicolor (abaxial faces silvery, adaxial green), lobes acute | → 25 |
23. Leaves silver-gray, lobes rounded (coastal dunes) | A. stelleriana |
23. Leaves bright green, lobes acute (not coastal dunes) | → 24 |
24. Involucres (4–)5–8 × 4–10 mm; 0–3800 m | A. norvegica |
24. Involucres 2.5–3.5 × 2–4.5 mm; 100–2400 m | A. packardiae |
25. Plants 30–100 cm, lemon-scented; heads usually erect (subalpine and alpine). | A. michauxiana |
25. Plants 15–70 cm, not lemon-scented; heads usually nodding | → 26 |
26. Perennials (widely spreading, stems brittle); garden escapes, c, e North America. | A. pontica |
26. Biennials or perennials (erect) | → 27 |
27. Leaves pinnatifid (lobes 3–5, 0.5–1 mm wide); grasslands or deserts, 600– 2900 m | A. carruthii |
27. Leaves 2–3-pinnately lobed (lobes elliptic, 2–6 mm wide); w mountains, 2200–3100 m | A. franserioides |
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FNA vol. 19, p. 532. |
FNA vol. 19, p. 520. |
Asteraceae > tribe Anthemideae > Artemisia > subg. Artemisia |
Asteraceae > tribe Anthemideae > Artemisia |
A. abrotanum, A. absinthium, A. alaskana, A. aleutica, A. annua, A. arbuscula, A. biennis, A. bigelovii, A. borealis, A. californica, A. campestris, A. cana, A. carruthii, A. douglasiana, A. dracunculus, A. filifolia, A. franserioides, A. frigida, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. nova, A. packardiae, A. palmeri, A. papposa, A. pattersonii, A. pedatifida, A. pontica, A. porteri, A. pycnocephala, A. pygmaea, A. rigida, A. rothrockii, A. rupestris, A. scopulorum, A. senjavinensis, A. serrata, A. spiciformis, A. suksdorfii, A. tilesii, A. tridentata, A. tripartita, A. vulgaris |
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A. abrotanum, A. alaskana, A. annua, A. biennis, A. californica, A. carruthii, A. douglasiana, A. franserioides, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. packardiae, A. palmeri, A. papposa, A. pontica, A. senjavinensis, A. serrata, A. stelleriana, A. suksdorfii, A. tilesii, A. vulgaris |
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A. section Abrotanum |
Besser: Nouv. Mém. Soc. Imp. Naturalistes Moscou 3: 79, plate 5. (1834) |
unknown |
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