Artemisia campestris |
Artemisia tilesii |
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beach wormwood, field sagewort, field wormwood, northern wormwood, Pacific sagewort, sand wormwood |
Aleutian mugwort, Cascade wormwood, Tilesius' wormwood |
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Habit | Biennials or perennials, (10–)30–80(–150) cm, faintly aromatic; taprooted, caudices branched. | Perennials, 20–60(–80) cm, mildly aromatic (rhizomes coarse). | ||||||||
Stems | usually 1–5, turning reddish brown, (often ribbed) tomentose or glabrous. |
1–3, erect, white, tomentose (on distal branches, hairs appressed) or glabrate. |
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Leaves | persistent or deciduous, mostly basal; basal blades 4–12 cm; cauline gradually reduced, 2–4 × 0.5–1.5 cm, 2–3-pinnately lobed, lobes linear to narrowly oblong, apices acute, faces densely to sparsely white-pubescent. |
basal and cauline, bicolor (white and green); blades (basal) linear to broadly lanceolate, 3–7(–10) × 2–5(–6) cm, coarsely pinnately lobed (cauline becoming linear distally), faces tomentose (abaxial) or glabrous (adaxial). |
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Involucres | broadly turbinate, 2.5–3(–5) × 2–3.5(–7) mm. |
broadly campanulate, 4–5 × 3.5–4 mm. |
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Florets | pistillate 5–20; functionally staminate 12–30; corollas pale yellow, sparsely hairy or glabrous. |
pistillate 9; bisexual 25–60; corollas yellow, 1.5–3 mm, glabrous (style branches included, erect, linear, relatively short, short-ciliate). |
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Phyllaries | (margins scarious) glabrous or villous-tomentose. |
(violet-brown) oval (outer) to elliptic or lanceolate, sparsely tomentose. |
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Heads | (pedunculate) in (mostly leafless) paniculiform arrays. |
in compact to broadly branched, paniculiform arrays 1–20 × 2–6 cm. |
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Cypselae | oblong-lanceoloid, somewhat compressed, 0.8–1 mm, faintly nerved, glabrous. |
oblong-linear (angular), 1.2–1.5 mm, glabrous. |
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2n | = 18, 36. |
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Artemisia campestris |
Artemisia tilesii |
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Phenology | Flowering mid summer–early fall. | |||||||||
Habitat | Arctic and alpine tundra, sandy, rocky slopes near shorelines | |||||||||
Elevation | 0–2000 m (0–6600 ft) | |||||||||
Distribution |
AK; AR; AZ; CA; CO; CT; FL; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NU; ON; QC; SK; especially mountains and high latitudes; Eurasia
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AK; ID; MT; OR; WA; AB; BC; MB; NT; NU; ON; QC; SK; YT; Asia (Russia)
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Discussion | Subspecies ca. 7 (3 in the flora). Artemisia campestris varies; each morphologic form grades into another. The present circumscription is conservative in that only three subspecies are recognized; the subspecies usually can be separated geographically as well as morphologically. Populations in western North America consist primarily of subsp. pacifica; east of the continental divide, plants are assigned to subsp. canadensis in northern latitudes and to subsp. caudata in southern latitudes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Artemisia tilesii has a bewildering array of variation in leaf and inflorescence morphology that has been separated into four infraspecific taxa recognized in some floras. I am unable to separate these taxa consistently and am including them within a broad circumscription of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 506. | FNA vol. 19, p. 533. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | A. hookeriana, A. hultenii, A. tilesii var. aleutica, A. tilesii var. elatior, A. tilesii subsp. gormanii, A. tilesii subsp. hultenii, A. tilesii var. unalaschcensis, A. unalaskensis var. aleutica, A. vulgaris subsp. tilesii | |||||||||
Name authority | Linnaeus: Sp. Pl. 2: 846. (1753) | Ledebour: Mém. Acad. Imp. Sci. St. Pétersbourg Hist. Acad. 5: 568. (1814) | ||||||||
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