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beach wormwood, field sagewort, field wormwood, northern wormwood, Pacific sagewort, sand wormwood

Aleutian mugwort, Cascade wormwood, Tilesius' wormwood

Habit Biennials or perennials, (10–)30–80(–150) cm, faintly aromatic; taprooted, caudices branched. Perennials, 20–60(–80) cm, mildly aromatic (rhizomes coarse).
Stems

usually 1–5, turning reddish brown, (often ribbed) tomentose or glabrous.

1–3, erect, white, tomentose (on distal branches, hairs appressed) or glabrate.

Leaves

persistent or deciduous, mostly basal;

basal blades 4–12 cm;

cauline gradually reduced, 2–4 × 0.5–1.5 cm, 2–3-pinnately lobed, lobes linear to narrowly oblong, apices acute, faces densely to sparsely white-pubescent.

basal and cauline, bicolor (white and green);

blades (basal) linear to broadly lanceolate, 3–7(–10) × 2–5(–6) cm, coarsely pinnately lobed (cauline becoming linear distally), faces tomentose (abaxial) or glabrous (adaxial).

Involucres

broadly turbinate, 2.5–3(–5) × 2–3.5(–7) mm.

broadly campanulate, 4–5 × 3.5–4 mm.

Florets

pistillate 5–20; functionally staminate 12–30;

corollas pale yellow, sparsely hairy or glabrous.

pistillate 9;

bisexual 25–60;

corollas yellow, 1.5–3 mm, glabrous (style branches included, erect, linear, relatively short, short-ciliate).

Phyllaries

(margins scarious) glabrous or villous-tomentose.

(violet-brown) oval (outer) to elliptic or lanceolate, sparsely tomentose.

Heads

(pedunculate) in (mostly leafless) paniculiform arrays.

in compact to broadly branched, paniculiform arrays 1–20 × 2–6 cm.

Cypselae

oblong-lanceoloid, somewhat compressed, 0.8–1 mm, faintly nerved, glabrous.

oblong-linear (angular), 1.2–1.5 mm, glabrous.

2n

= 18, 36.

Artemisia campestris

Artemisia tilesii

Phenology Flowering mid summer–early fall.
Habitat Arctic and alpine tundra, sandy, rocky slopes near shorelines
Elevation 0–2000 m (0–6600 ft)
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; FL; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NU; ON; QC; SK; especially mountains and high latitudes; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; ID; MT; OR; WA; AB; BC; MB; NT; NU; ON; QC; SK; YT; Asia (Russia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies ca. 7 (3 in the flora).

Artemisia campestris varies; each morphologic form grades into another. The present circumscription is conservative in that only three subspecies are recognized; the subspecies usually can be separated geographically as well as morphologically. Populations in western North America consist primarily of subsp. pacifica; east of the continental divide, plants are assigned to subsp. canadensis in northern latitudes and to subsp. caudata in southern latitudes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Artemisia tilesii has a bewildering array of variation in leaf and inflorescence morphology that has been separated into four infraspecific taxa recognized in some floras. I am unable to separate these taxa consistently and am including them within a broad circumscription of the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials; stems 2–5; basal rosettes persistent
subsp. pacifica
1. Biennials; stems 1(–3); basal rosettes not persistent (withering before flowering)
→ 2
2. Involucres globose, 3–4 × 3.5–5 mm; n of 50°, primarily Canada
subsp. canadensis
2. Involucres turbinate, 2–3 × 2–3 mm; s of 50°, e from Rocky Mountains to coastal North America
subsp. caudata
Source FNA vol. 19, p. 506. FNA vol. 19, p. 533.
Parent taxa Asteraceae > tribe Anthemideae > Artemisia > subg. Drancunculus Asteraceae > tribe Anthemideae > Artemisia > subg. Artemisia
Sibling taxa
A. abrotanum, A. absinthium, A. alaskana, A. aleutica, A. annua, A. arbuscula, A. biennis, A. bigelovii, A. borealis, A. californica, A. cana, A. carruthii, A. douglasiana, A. dracunculus, A. filifolia, A. franserioides, A. frigida, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. nova, A. packardiae, A. palmeri, A. papposa, A. pattersonii, A. pedatifida, A. pontica, A. porteri, A. pycnocephala, A. pygmaea, A. rigida, A. rothrockii, A. rupestris, A. scopulorum, A. senjavinensis, A. serrata, A. spiciformis, A. stelleriana, A. suksdorfii, A. tilesii, A. tridentata, A. tripartita, A. vulgaris
A. abrotanum, A. absinthium, A. alaskana, A. aleutica, A. annua, A. arbuscula, A. biennis, A. bigelovii, A. borealis, A. californica, A. campestris, A. cana, A. carruthii, A. douglasiana, A. dracunculus, A. filifolia, A. franserioides, A. frigida, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. nova, A. packardiae, A. palmeri, A. papposa, A. pattersonii, A. pedatifida, A. pontica, A. porteri, A. pycnocephala, A. pygmaea, A. rigida, A. rothrockii, A. rupestris, A. scopulorum, A. senjavinensis, A. serrata, A. spiciformis, A. stelleriana, A. suksdorfii, A. tridentata, A. tripartita, A. vulgaris
Subordinate taxa
A. campestris subsp. canadensis, A. campestris subsp. caudata, A. campestris subsp. pacifica
Synonyms A. hookeriana, A. hultenii, A. tilesii var. aleutica, A. tilesii var. elatior, A. tilesii subsp. gormanii, A. tilesii subsp. hultenii, A. tilesii var. unalaschcensis, A. unalaskensis var. aleutica, A. vulgaris subsp. tilesii
Name authority Linnaeus: Sp. Pl. 2: 846. (1753) Ledebour: Mém. Acad. Imp. Sci. St. Pétersbourg Hist. Acad. 5: 568. (1814)
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