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black sagebrush, dwarf sagebrush, little sagebrush, low sagebrush

armoise de Steller, beach wormwood, dusty miller, oldwoman, Steller's wormwood

Habit Shrubs, 10–30(–50) cm, aromatic; root-sprouting. Perennials, (15–)20–60(–70) cm (mat-forming), sometimes faintly aromatic (rhizomes creeping, relatively thin).
Stems

gray-green to brown, glabrate (diffusely branched from bases, brittle).

1–3, erect or ascending, white, simple (stout), densely tomentose to floccose.

Leaves

(vegetative stems) persistent, gray-green;

blades broadly to narrowly cuneate, 3–10 × 2–5 mm, lobed (lobes 3, oblong-linear, to 1/3 blade lengths, mostly 1–3 mm wide, flat, obtuse, laterals sometimes 2–3-fid; leaves on flowering stems deciduous, blades narrowly cuneate, deeply 3-lobed), faces densely hairy (not sticky resinous).

basal and cauline (petiolate), silver-gray;

blades oblanceolate, (proximalmost) 3–10 × 1–5 cm, pinnatifid (lobes relatively broad, rounded; distal leaves, on flowering stems, smaller), faces densely tomentose.

Involucres

campanulate or globose-ovoid, (1.5–)2–4(–5) × 1.5–4.5 mm.

broadly campanulate, 5–8 × 6–7 mm.

Florets

4–6(–10);

corollas 1.5–2 mm, glabrous.

pistillate 12–16;

bisexual 25–30;

corollas yellow (narrow or tubular), 3.2–4 mm (unusually large), glabrous or sparsely hairy (style branches prominent, erect, blunt).

Phyllaries

(margins green) ovate (outer) to oblong, pubescent or tomentose.

broadly lanceolate, tomentose.

Heads

usually borne singly, rarely (1–4, erect, mostly sessile, in pedunculate clusters) in spiciform or paniculiform arrays 2–9 × 0.5–2 cm (branches slender).

(erect or spreading, peduncles 0 or to 3 mm) in dense, paniculiform, racemiform, or spiciform arrays 8–20 × 2–4 cm.

Cypselae

(light brown) 0.7–0.8 mm, resinous.

(dark brown) narrowly oblong-linear (slightly flattened, smooth), 3–4 mm, glabrous.

2n

= 18.

Artemisia arbuscula

Artemisia stelleriana

Phenology Flowering early spring–fall.
Habitat Sandy soils, coastal strand
Elevation 0–200 m (0–700 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CT; DE; FL; LA; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; VA; VT; WA; WI; WV; NB; NF; NS; ON; PE; QC; SPM; n Europe; e Asia (Japan, Kamchatka)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Artemisia arbuscula is one of the more perplexing species in the Tridentatae complex. Anatomic and morphologic characteristics suggest multiple hybrid origins for the subspecies. Deciduous leaves of flowering stems in plants that otherwise have persistent leaves suggest a hybrid origin involving plants of the A. tridentata and A. cana lineages. In most instances, populations of A. arbuscula appear to be reproductively stable. The disposition of Artemisia arbuscula subsp. longicaulis Winward & McArthur (with 2n = 54) has not been determined.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Artemisia stelleriana is apparently native along the western tip of the Aleutian islands (D. F. Murray, pers. comm.). It is an attractive ornamental and, in parts of its range in the flora area, it appears to have escaped from cultivation and is naturalized in beach dunes and other sandy habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres 2–4.5 mm diam.; usually in rocky soils; flowering mid–late summer
subsp. arbuscula
1. Involucres 1.5–2.5 mm diam.; clays or stony soils; flowering early spring–late summer
→ 2
2. Leaves broadly cuneate (4–10 × 2–5 mm, often irregularly lobed, lobes rounded, middle lobes overlapping lateral lobes); usually in clay soils; flower- ing early spring
subsp. longiloba
2. Leaves narrowly cuneate (5–10 × 3–6 mm, lobed, lobes 1/2+ blade lengths, laterals to 1 mm wide, of-ten acute); usually in stony soils; flowering mid–late summer
subsp. thermopola
Source FNA vol. 19, p. 511. FNA vol. 19, p. 532.
Parent taxa Asteraceae > tribe Anthemideae > Artemisia > subg. Tridentatae Asteraceae > tribe Anthemideae > Artemisia > subg. Artemisia
Sibling taxa
A. abrotanum, A. absinthium, A. alaskana, A. aleutica, A. annua, A. biennis, A. bigelovii, A. borealis, A. californica, A. campestris, A. cana, A. carruthii, A. douglasiana, A. dracunculus, A. filifolia, A. franserioides, A. frigida, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. nova, A. packardiae, A. palmeri, A. papposa, A. pattersonii, A. pedatifida, A. pontica, A. porteri, A. pycnocephala, A. pygmaea, A. rigida, A. rothrockii, A. rupestris, A. scopulorum, A. senjavinensis, A. serrata, A. spiciformis, A. stelleriana, A. suksdorfii, A. tilesii, A. tridentata, A. tripartita, A. vulgaris
A. abrotanum, A. absinthium, A. alaskana, A. aleutica, A. annua, A. arbuscula, A. biennis, A. bigelovii, A. borealis, A. californica, A. campestris, A. cana, A. carruthii, A. douglasiana, A. dracunculus, A. filifolia, A. franserioides, A. frigida, A. furcata, A. globularia, A. glomerata, A. laciniata, A. longifolia, A. ludoviciana, A. michauxiana, A. nesiotica, A. norvegica, A. nova, A. packardiae, A. palmeri, A. papposa, A. pattersonii, A. pedatifida, A. pontica, A. porteri, A. pycnocephala, A. pygmaea, A. rigida, A. rothrockii, A. rupestris, A. scopulorum, A. senjavinensis, A. serrata, A. spiciformis, A. suksdorfii, A. tilesii, A. tridentata, A. tripartita, A. vulgaris
Subordinate taxa
A. arbuscula subsp. arbuscula, A. arbuscula subsp. longiloba, A. arbuscula subsp. thermopola
Synonyms A. tridentata subsp. arbuscula, A. tridentata var. arbuscula, Seriphidium arbusculum
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 398. (1841) Besser: Nouv. Mém. Soc. Imp. Naturalistes Moscou 3: 79, plate 5. (1834)
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