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broad-leaf arnica, daffodil leopardbane, mountain arnica

arnica

Habit Plants 10–50 cm. Perennials, 5–100 cm (rhizomes relatively long and thin; caudices woody, relatively short and thick).
Stems

usually simple, sometimes branched distally.

erect, simple or branched.

Leaves

2–4(–6) pairs, cauline (basal leaves often persistent on sterile rosettes), petiolate (proximal, petioles relatively short, broadly winged) or sessile (mid and distal);

blades lance-elliptic to ovate, 2–10 × 1–6 cm, margins serrate to dentate, apices obtuse to acute, faces glabrous or sparsely villous.

basal (sterile basal rosettes often present) and/or cauline; mostly opposite (usually 1–10 pairs, distalmost sometimes alternate and usually smaller); petiolate or sessile;

blades mostly cordate, deltate, elliptic, lanceolate, linear, oblanceolate, oblong, obovate, ovate, or spatulate, margins entire or toothed (usually dentate, denticulate, or serrate, sometimes crenate or slightly lobed), faces glabrous, hirsute, hispidulous, pilose, puberulent, scabrous, tomentose, villous, or woolly, often stipitate-glandular as well.

Involucres

narrowly turbinate.

campanulate, hemispheric, or turbinate, mostly 6–20+ mm diam.

Receptacles

convex, smooth or pitted, epaleate.

Ray florets

8–15;

corollas yellow.

0, or 5–22, pistillate, fertile;

corollas yellow to orange.

Disc florets

20–90;

corollas yellow;

anthers yellow.

10–120, usually bisexual and fertile (functionally staminate in A. dealbata);

corollas usually yellow, rarely cream, tubes shorter than funnelform throats, lobes 5, ± deltate (anthers usually yellow, purple in A. lessingii and A. unalaschcensis).

Phyllaries

8–20, lanceolate to oblanceolate.

persistent, 5–23 in (1–)2 series.

Heads

1 or 3–5(–9).

radiate or discoid, borne singly or in cymiform or corymbiform arrays.

Cypselae

dark brown, 5–9 mm, sparsely villous;

pappi white, bristles barbellate.

(gray or brown to black) ± conic, fusiform, or obovoid-cylindric, nerves 5–10(–20), faces hairy, glandular, or glabrous;

pappi usually persistent, of 10–50 white or stramineous to tawny, fine, barbellate or subplumose to plumose bristles (0 in A. dealbata).

x

= 19.

2n

= 38, 76.

Arnica latifolia

Arnica

Phenology Flowering Jul–Aug.
Habitat Relatively moist, montane conifer forests to subalpine meadows
Elevation 500–3300 m (1600–10800 ft)
Distribution
from FNA
AK; CA; CO; ID; MT; OR; UT; WA; WY; AB; BC; NT; YT
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Mexico; Europe; Asia (Japan, Russia)
[BONAP county map]
Discussion

Species 29 (26 in the flora).

Arnica is circumboreal, predominantly montane, and exhibits maximum species diversity in western North America. It includes common and very widespread species as well as relatively uncommon, narrow endemics. Polyploidy and apomixis are common in the genus, resulting in considerable morphologic variability. Arnica montana from Europe has been used medicinally for centuries, and unsubstantiated claims have been made regarding the medicinal properties of some North American species.

Here, in key leads and descriptions, numbers of pairs of leaves refer to flowering shoots.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pappi 0 (ray corollas marcescent; disc florets functionally staminate)
A. dealbata
1. Pappi of 10–50, fine bristles
→ 2
2. Anthers purple
→ 3
2. Anthers yellow
→ 4
3. Leaves mostly cauline; heads erect; pappus bristles subplumose
A. unalaschcensis
3. Leaves mostly basal; heads nodding; pappus bristles usually barbellate
A. lessingii
4. Ray florets 0 (disc corollas sometimes dilated in A. parryi, resembling rays)
→ 5
4. Ray florets 5–22
→ 9
5. Cauline leaves usually sessile, rarely broadly petiolate
→ 6
5. Cauline leaves petiolate (petioles sometimes relatively broad)
→ 7
6. Leaves: (reticulate venation inconspicuous) margins ± entire, abaxial and adaxial faces sparsely to densely pilose and stipitate-glandular; disc corollascream
A. viscosa
6. Leaves: (reticulate venation conspicuous) margins irregularly and coarsely serrate, abaxial faces pilose and stipitate-glandular, adaxial glabrate tostipitate-glandular; disc corollas yellow
A. venosa
7. Heads 1–14 (nodding in bud); phyllaries linear to narrowly lanceolate; pappi usually stramineous, rarely tawny, bristles barbellate to ± subplumose
A. parryi
7. Heads 1–30 (erect); phyllaries lanceolate, ovate, or ovate-lanceolate; pappi white, bristles usually barbellate, sometimes subplumose
→ 8
8. Leaves lanceolate, ovate, or subcordate (petioles usually narrow or scarcely winged); cypselae stipitate-glandular and hirsute (hairs duplex)
A. discoidea
8. Leaves elliptic-ovate to spatulate (petioles relatively broad); cypselae sparsely stipitate-glandular (lacking duplex hairs)
A. spathulata
9. Leaves mostly basal (blades usually with 3, 5, or 7 prominent, subparallel veins; cauline leaves 0 or 1–2 and much reduced)
→ 10
9. Leaves mostly cauline, sometimes also basal (blades lacking prominent, subparallel veins)
→ 12
10. Leaves broadly elliptic, ovate, or rhombic; involucres campanulate; raysyellow; e United States
A. acaulis
10. Leaves oblanceolate, oblong, oval, or spatulate; involucres hemispheric; rays yellow-orange; Canada, w United States
→ 11
11. Leaf blades 4.5–20 × 0.5–2.5 cm (axils with tufts of brown wool)
A. fulgens
11. Leaf blades 3.5–14.5 × 0.6–2.4 cm (axils lacking tufts of brown wool)
A. sororia
12. Cauline leaves (at least at mid stems and exclusive of distal reduced leaves) sessile
→ 13
12. Cauline leaves (at least mid-stem pair) petiolate (petioles sometimes broad)
→ 18
13. Pappi white, bristles barbellate
→ 14
13. Pappi stramineous to tawny, bristles usually subplumose to plumose sometimes barbellate
→ 15
14. Leaf blades ovate to elliptic-lanceolate, 2–10 × 1–6 cm, margins serrate to dentate
A. latifolia
14. Leaf blades oblanceolate to spatulate, 2–7 × 0.5–2.5 cm, margins usually entire, sometimes denticulate
A. rydbergii
15. Cauline leaves (2–)3(–4) pairs (pappus bristles with deep, amberlikedeposits)
A. mollis
15. Cauline leaves usually 4–10 pairs (pappus bristles lacking deep, amberlike deposits)
→ 16
16. Phyllaries 8–23, lanceolate to linear (apices with conspicuoustufts of white hairs)
A. chamissonis
16. Phyllaries 8–20, narrowly to broadly lanceolate (apices lacking conspicuous tufts of white hairs)
→ 17
17. Leaves lance-elliptic, obovate, or ovate, margins subentire todentate-serrate (basal leaves present at flowering)
A. lanceolata
17. Leaves lanceolate to lance-elliptic, margins usually entire(basal leaves withered by flowering)
A. longifolia
18. Cauline leaf blades elliptic, lanceolate, linear, oblanceolate, ovate, or spatulate (lengths 2.5–10 times widths; basal leaves withered by flowering)
→ 19
18. Cauline leaf blades cordate, deltate, elliptic, ovate, ovate-lanceolate, or subcordate (lengths 1–3 times widths; basal leaves present at flowering)
→ 22
19. Heads 1–8 (erect)
→ 20
19. Heads 1–3 (usually nodding)
→ 21
20. Leaves 3–7 pairs (petioles relatively narrow, lengths ± equaling blades)
A. lonchophylla
20. Leaves 1–5 pairs (petioles relatively broadly to narrowly winged, lengths shorterthan blades)
A. angustifolia
21. Leaves: margins usually entire, rarely denticulate or slightly undulate, faces ± densely stipitate-glandular; cypselae ± glandular; Alberta
A. louiseana
21. Leaves: margins usually denticulate to dentate, rarely entire, faces rarely stipitate-glandular; cypselae rarely glandular; e Canada or Alaska, nw Canada (not Alberta)
A. griscomii
22. Pappi stramineous to tawny, bristles subplumose (sterile basal leaf rosetteslacking)
A. ovata
22. Pappi white, bristles usually barbellate, rarely subplumose (sterile basal leaf rosettes present)
→ 23
23. Leaves (relatively thick, often succulent and reddish) glabrous or scabrous; heads 1(–3; often nodding in bud; serpentine in nw California, sw Oregon)
A. cernua
23. Leaves (relatively thin, not succulent nor reddish) hairy; heads 1 or 3–15 (erect in bud; plants not of serpentine)
→ 24
24. Stems usually much branched distally (often in dense clumps); heads (1–)5–15; cypselae black
A. gracilis
24. Stems usually simple (not in dense clumps); heads 1 or 3–5(–10); cypselae gray
→ 25
25. Leaves cordate, ovate, or subcordate, margins dentate; involucres broadlycampanulate
A. cordifolia
25. Leaves elliptic to ovate, margins entire or denticulate; involucres campanulate-turbinate
A. nevadensis
Source FNA vol. 21, p. 376. FNA vol. 21, p. 366. Author: Steven J. Wolf.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Arnica Asteraceae > tribe Heliantheae > subtribe Chaenactidinae
Sibling taxa
A. acaulis, A. angustifolia, A. cernua, A. chamissonis, A. cordifolia, A. dealbata, A. discoidea, A. fulgens, A. gracilis, A. griscomii, A. lanceolata, A. lessingii, A. lonchophylla, A. longifolia, A. louiseana, A. mollis, A. nevadensis, A. ovata, A. parryi, A. rydbergii, A. sororia, A. spathulata, A. unalaschcensis, A. venosa, A. viscosa
Subordinate taxa
A. acaulis, A. angustifolia, A. cernua, A. chamissonis, A. cordifolia, A. dealbata, A. discoidea, A. fulgens, A. gracilis, A. griscomii, A. lanceolata, A. latifolia, A. lessingii, A. lonchophylla, A. longifolia, A. louiseana, A. mollis, A. nevadensis, A. ovata, A. parryi, A. rydbergii, A. sororia, A. spathulata, A. unalaschcensis, A. venosa, A. viscosa
Name authority Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 147. (1832) Linnaeus: Sp. Pl. 2: 884. (1753): Gen. Pl. ed. 5, 376. (1754)
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