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pineland threeawn, wiregrass

Habit Plants perennial; cespitose, occasionally with rhizomes. Plants annual or perennial; usually cespitose.
Culms

60-120 cm, stiffly erect, unbranched.

annual, erect, solid or hollow, usually unbranched.

Leaves

basal or nearly basal;

sheaths shorter than the internodes, smooth and mostly glabrous abaxially, sometimes with a line of widely spaced, 0.5-1 mm hairs over the midvein, not disintegrating into threadlike fibers at maturity;

collars glabrous, sometimes with a few conspicuous hairs at the sides;

ligules 0.1-0.3 mm;

blades 15-50 cm long, 0.3-1 mm wide, tightly involute, stiff, yellow-green, abaxial surfaces villous on both sides of the midvein, at least on the basal portion, hairs 0.6-1.5 mm, adaxial surfaces densely scabrous or densely short pubescent.

distichous;

sheaths usually open;

auricles absent;

abaxial ligules absent or of hairs;

adaxial ligules membranous and ciliate or of hairs;

blades without pseudopetioles;

mesophyll cells radiate or non-radiate;

adaxial palisade layer absent;

fusoid cells absent;

arm cells absent;

kranz anatomy absent or present, when present, with 1 or 2 parenchyma sheaths;

midribs simple;

adaxial bulliform cells present;

stomatal subsidiary cells dome-shaped or triangular;

bicellular microhairs present, with long, slender, thin-walled terminal cells.

Inflorescences

paniculate, 20-35 cm, 5-8 mm wide;

nodes glabrous;

primary branches 2-5 cm, appressed, without axillary pulvini, with 4-12 spikelets, spikelet-bearing to the base.

terminal, not leafy, usually panicles, sometimes spikes or racemes;

disarticulation above the glumes.

Spikelets

appressed.

bisexual, with 1 floret;

rachilla extension absent.

Glumes

subequal, glabrous, light brown or tan, usually 1(2)-veined, bifid and awned, awns 1.5-2.5 mm;

lower glumes 7-10 mm;

upper glumes 6-9 mm;

calluses 0.4-0.6 mm;

lemmas 6-9 mm, glabrous, light-colored when young, reddish when mature, column 1-2 mm, not twisted, junction with the awns inconspicuous;

awns (7)10-15(22) mm, subequal, usually horizontally spreading or curving downward, not disarticulating at maturity;

anthers 3, about 3 mm, reddish-brown.

2, usually longer than the florets, usually acute or acuminate;

florets terete or laterally compressed, with well-developed calluses;

lemmas 1- or 3-veined, more or less coriaceous, with a germination flap, lemma margins overlapping at maturity and concealing the paleas, apices evidently 3-awned;

awn bases often forming a column, lateral awns occasionally reduced or absent;

paleas less than 1/2 as long as the lemmas;

lodicules usually present, 2, free, membranous, glabrous, heavily vascularized;

anthers 1-3;

ovaries glabrous;

haustorial synergids absent;

styles 2, free to the base but close.

Caryopses

4-5 mm, chestnut brown.

usually fusiform, falling with the lemma and palea attached;

hila short or long, linear;

endosperm hard, without lipid;

starch grains compound;

embryos small or large relative to the caryopses;

epiblasts absent;

scutellar cleft present or absent;

mesocotyl internode elongated;

embryonic leaf margins meeting, x = 11, 12.

2n

= unknown.

Aristida stricta

Poaceae subfam. aristidoideae

Distribution
from FNA
NC; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

Aristida stricta grows in pine barrens and sandy fields of the coastal plain from Louisiana to North Carolina. Peet (1993) segregated northern populations of A. stricta as a separate species, A. beyrichiana Trin. & Rupr., based on pubescence patterns of the sheath and blades. Investigations into alloyzyme diversity (Walters et al. 1994), anatomy, morphology, and phenotypic expression (garden transplants) led Kesler (2000) to conclude that such a segregation was not justified; pubescence patterns particularly were inconclusive. Consequently, A. beyrichiana is treated here as part of A. stricta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The subfamily Aristidoideae includes only one tribe, the Aristideae. Other taxonomists have generally included the Aristidoideae, with the Danthonieae and Arundineae, in the Arundinoideae (e.g., Watson et al. 1985; Clayton and Renvoize 1986; Kellogg and Campbell 1987), but Esen and Hilu (1991) demonstrated that Aristida is clearly distinct from the Danthonieae and Arundineae in terms of its prolamins. Subsequent work has provided further support for the monophyly of the three tribes, but their position relative to each other and other members of the PACCAD clade is more equivocal.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 335. FNA vol. 25, p. 314. Author: Grass Phylogeny Working Group; Kelly W. Allred;.
Parent taxa Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida Poaceae
Sibling taxa
A. adscensionis, A. arizonica, A. basiramea, A. californica, A. condensata, A. desmantha, A. dichotoma, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurascens, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. ternipes, A. tuberculosa
Subordinate taxa
Synonyms A. beyrichiana
Name authority Michx. Caro
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