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pineland threeawn, wiregrass

arrow-feather threeawn

Habit Plants perennial; cespitose, occasionally with rhizomes. Plants perennial; cespitose, bases knotty, without rhizomes.
Culms

60-120 cm, stiffly erect, unbranched.

40-100 cm tall, 1-4 mm thick at the base, erect, branching at the base, shoots becoming thickened and somewhat fan-shaped upwards.

Leaves

basal or nearly basal;

sheaths shorter than the internodes, smooth and mostly glabrous abaxially, sometimes with a line of widely spaced, 0.5-1 mm hairs over the midvein, not disintegrating into threadlike fibers at maturity;

collars glabrous, sometimes with a few conspicuous hairs at the sides;

ligules 0.1-0.3 mm;

blades 15-50 cm long, 0.3-1 mm wide, tightly involute, stiff, yellow-green, abaxial surfaces villous on both sides of the midvein, at least on the basal portion, hairs 0.6-1.5 mm, adaxial surfaces densely scabrous or densely short pubescent.

cauline;

sheaths mostly longer than the internode, mostly or completely glabrous, sometimes pilose, particularly along the margins and at the throat, remaining intact at maturity;

collars glabrous or pilose;

ligules about 0.2 mm;

blades 10-25 cm long, 1-3 mm wide, usually flat, usually lax, sometimes sinuous to curling at maturity, glabrous, pale green, drying brownish.

Inflorescences

paniculate, 20-35 cm, 5-8 mm wide;

nodes glabrous;

primary branches 2-5 cm, appressed, without axillary pulvini, with 4-12 spikelets, spikelet-bearing to the base.

paniculate, (15)20-55 cm long, 0.5-2(3) cm wide, often nodding;

nodes glabrous, sometimes scabrous, lower nodes usually associated with more than 2 spikelets;

primary branches 1-5 cm, tightly appressed to loosely ascending, without axillary pulvini, with 1-8 spikelets.

Spikelets

appressed.

appressed.

Glumes

subequal, glabrous, light brown or tan, usually 1(2)-veined, bifid and awned, awns 1.5-2.5 mm;

lower glumes 7-10 mm;

upper glumes 6-9 mm;

calluses 0.4-0.6 mm;

lemmas 6-9 mm, glabrous, light-colored when young, reddish when mature, column 1-2 mm, not twisted, junction with the awns inconspicuous;

awns (7)10-15(22) mm, subequal, usually horizontally spreading or curving downward, not disarticulating at maturity;

anthers 3, about 3 mm, reddish-brown.

5-10 mm, lower glumes from 3/4 as long as to 1-4 mm longer than the upper glumes, glabrous or sparsely appressed-pubescent, 1-2-veined, 1-keeled, tan to purplish, unawned or the awns no longer than 1 mm;

calluses 0.4-0.8 mm;

lemmas 4-8 mm, glabrous, mostly light tan or gray, often spotted or banded, beak not twisted, junction with the awns not evident;

awns 8-25 mm, equal or subequal in length, curved, arcuate, or spirally coiled at the base, not disarticulating at maturity;

central awns sometimes thicker than the lateral awns, erect to arcuate-reflexed;

lateral awns straight and erect, ascending, or divergent;

anthers 1 or 3, 1-1.5 mm, brown.

Caryopses

4-5 mm, chestnut brown.

3-5 mm, chestnut brown.

2n

= unknown.

= unknown.

Aristida stricta

Aristida purpurascens

Distribution
from FNA
NC; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Aristida stricta grows in pine barrens and sandy fields of the coastal plain from Louisiana to North Carolina. Peet (1993) segregated northern populations of A. stricta as a separate species, A. beyrichiana Trin. & Rupr., based on pubescence patterns of the sheath and blades. Investigations into alloyzyme diversity (Walters et al. 1994), anatomy, morphology, and phenotypic expression (garden transplants) led Kesler (2000) to conclude that such a segregation was not justified; pubescence patterns particularly were inconclusive. Consequently, A. beyrichiana is treated here as part of A. stricta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central awns divaricate to reflexed, about twice as thick at the base as the lateral awns
var. virgata
1. Central and lateral awns divergent, all about the same thickness at the base.
→ 2
2. Lower glumes usually longer than the upper glumes; awns straight or only slightly contorted at the base; blades 1-3 mm wide, often curling
var. purpurascens
2. Lower glumes shorter than or equal to the upper glumes; awns spirally contorted at the base; blades mostly about 1 mm wide, usually not curling
var. tenuispica
Source FNA vol. 25, p. 335. FNA vol. 25, p. 338.
Parent taxa Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida
Sibling taxa
A. adscensionis, A. arizonica, A. basiramea, A. californica, A. condensata, A. desmantha, A. dichotoma, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurascens, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. ternipes, A. tuberculosa
A. adscensionis, A. arizonica, A. basiramea, A. californica, A. condensata, A. desmantha, A. dichotoma, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. stricta, A. ternipes, A. tuberculosa
Subordinate taxa
A. purpurascens var. purpurascens, A. purpurascens var. tenuispica, A. purpurascens var. virgata
Synonyms A. beyrichiana
Name authority Michx. Poir.
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