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pineland threeawn, wiregrass

churchmouse three-awn

Habit Plants perennial; cespitose, occasionally with rhizomes. Plants annual.
Culms

60-120 cm, stiffly erect, unbranched.

15-60 cm, erect or geniculate at the base, branching at most of the nodes.

Leaves

basal or nearly basal;

sheaths shorter than the internodes, smooth and mostly glabrous abaxially, sometimes with a line of widely spaced, 0.5-1 mm hairs over the midvein, not disintegrating into threadlike fibers at maturity;

collars glabrous, sometimes with a few conspicuous hairs at the sides;

ligules 0.1-0.3 mm;

blades 15-50 cm long, 0.3-1 mm wide, tightly involute, stiff, yellow-green, abaxial surfaces villous on both sides of the midvein, at least on the basal portion, hairs 0.6-1.5 mm, adaxial surfaces densely scabrous or densely short pubescent.

cauline;

sheaths usually shorter than the internodes, glabrous or sparsely pilose;

collars glabrous;

ligules less than 0.5 mm;

blades 3-10 cm long, 1-2 mm wide, flat to folded basally, involute distally, scabridulous on both surfaces, occasionally sparsely pilose adaxially, light green.

Inflorescences

paniculate, 20-35 cm, 5-8 mm wide;

nodes glabrous;

primary branches 2-5 cm, appressed, without axillary pulvini, with 4-12 spikelets, spikelet-bearing to the base.

paniculate or racemose, 2-11 cm long, to 1 cm wide;

nodes glabrous or strigillose;

primary branches 1-2 cm, appressed, without axillary pulvini, with 1-2 spikelets.

Spikelets

appressed.

partly overlapping, often in pairs, 1 spikelet subsessile, the other pedicellate.

Glumes

subequal, glabrous, light brown or tan, usually 1(2)-veined, bifid and awned, awns 1.5-2.5 mm;

lower glumes 7-10 mm;

upper glumes 6-9 mm;

calluses 0.4-0.6 mm;

lemmas 6-9 mm, glabrous, light-colored when young, reddish when mature, column 1-2 mm, not twisted, junction with the awns inconspicuous;

awns (7)10-15(22) mm, subequal, usually horizontally spreading or curving downward, not disarticulating at maturity;

anthers 3, about 3 mm, reddish-brown.

1-veined, light gray to dark purplish or brownish;

lower glumes 3-8(10) mm, from 1/2 as long as the upper glumes to nearly equaling them;

upper glumes 4-13 mm;

calluses 0.3-0.5 mm;

lemmas 3-11 mm, light gray to purplish, frequently mottled, midveins scabrous, elsewhere glabrous, scabridulous, or sparsely appressed-puberulent, junction with the awns not evident;

central awns 3-8 mm, coiled at the base, spreading distally;

lateral awns 1-4 mm, straight, erect;

anthers 3 and 2-3 mm, or 1 and about 0.25 mm.

Caryopses

4-5 mm, chestnut brown.

light brown.

2n

= unknown.

= unknown.

Aristida stricta

Aristida dichotoma

Distribution
from FNA
NC; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Aristida stricta grows in pine barrens and sandy fields of the coastal plain from Louisiana to North Carolina. Peet (1993) segregated northern populations of A. stricta as a separate species, A. beyrichiana Trin. & Rupr., based on pubescence patterns of the sheath and blades. Investigations into alloyzyme diversity (Walters et al. 1994), anatomy, morphology, and phenotypic expression (garden transplants) led Kesler (2000) to conclude that such a segregation was not justified; pubescence patterns particularly were inconclusive. Consequently, A. beyrichiana is treated here as part of A. stricta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Aristida dichotoma grows in sandy fields and clearings, disturbed sites and sterile ground, pine woods, and on granitic outcrops of the United States and southern Ontario. The two varieties have similar ecological preferences and extensive overlap in their ranges, but var. curtissii is somewhat more western in its distribution.

Aristida dichotoma is similar to A. basiramea, differing in its shorter lateral awns. Further study may show that the two should be treated as conspecific varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Glumes unequal; lemmas smooth or scabridulous, 6-11 mm long
var. curtissii
1. Glumes equal or subequal; lemmas sparsely appressed-pubescent, 3-8 mm long
var. dichotoma
Source FNA vol. 25, p. 335. FNA vol. 25, p. 328.
Parent taxa Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida
Sibling taxa
A. adscensionis, A. arizonica, A. basiramea, A. californica, A. condensata, A. desmantha, A. dichotoma, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurascens, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. ternipes, A. tuberculosa
A. adscensionis, A. arizonica, A. basiramea, A. californica, A. condensata, A. desmantha, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurascens, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. stricta, A. ternipes, A. tuberculosa
Subordinate taxa
A. dichotoma var. curtissii, A. dichotoma var. dichotoma
Synonyms A. beyrichiana
Name authority Michx. Michx.
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