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California three-awn, Mojave three-awn

Habit Plants perennial; sometimes flowering the first year. Plants annual or perennial; usually cespitose.
Culms

10-40 cm, highly branched above the base in age;

internodes glabrous or pubescent, sometimes nearly lanose.

annual, erect, solid or hollow, usually unbranched.

Leaves

cauline;

sheaths shorter than the internodes, glabrous or puberulent;

collars glabrous or pubescent at the sides;

ligules 0.5-1 mm;

blades usually less than 6 cm long, 0.5-1 mm wide, pale green, involute, glabrous or puberulent abaxially.

distichous;

sheaths usually open;

auricles absent;

abaxial ligules absent or of hairs;

adaxial ligules membranous and ciliate or of hairs;

blades without pseudopetioles;

mesophyll cells radiate or non-radiate;

adaxial palisade layer absent;

fusoid cells absent;

arm cells absent;

kranz anatomy absent or present, when present, with 1 or 2 parenchyma sheaths;

midribs simple;

adaxial bulliform cells present;

stomatal subsidiary cells dome-shaped or triangular;

bicellular microhairs present, with long, slender, thin-walled terminal cells.

Inflorescences

paniculate or racemose, 5-10 cm long, 1-2 cm wide, with few spikelets;

rachis nodes glabrous or with straight hairs;

primary branches 1-2 cm, appressed, without axillary pulvini.

terminal, not leafy, usually panicles, sometimes spikes or racemes;

disarticulation above the glumes.

Spikelets

appressed.

bisexual, with 1 floret;

rachilla extension absent.

Glumes

unequal, 1-2-veined;

lower glumes 4-10 mm;

upper glumes 7-15 mm;

calluses about 1 mm;

lemmas 5-7 mm, purple or mottled, junction of the lemma and awns evident;

awns twisted together basally into a 4-26 mm column, free portions 12-50 mm, those of the central and lateral awns similar in length, curved to arcuate basally, straight and divergent distally, disarticulating at the base of the column at maturity;

anthers 3, about 2 mm long.

2, usually longer than the florets, usually acute or acuminate;

florets terete or laterally compressed, with well-developed calluses;

lemmas 1- or 3-veined, more or less coriaceous, with a germination flap, lemma margins overlapping at maturity and concealing the paleas, apices evidently 3-awned;

awn bases often forming a column, lateral awns occasionally reduced or absent;

paleas less than 1/2 as long as the lemmas;

lodicules usually present, 2, free, membranous, glabrous, heavily vascularized;

anthers 1-3;

ovaries glabrous;

haustorial synergids absent;

styles 2, free to the base but close.

Caryopses

usually fusiform, falling with the lemma and palea attached;

hila short or long, linear;

endosperm hard, without lipid;

starch grains compound;

embryos small or large relative to the caryopses;

epiblasts absent;

scutellar cleft present or absent;

mesocotyl internode elongated;

embryonic leaf margins meeting, x = 11, 12.

2n

= 22.

Aristida californica

Poaceae subfam. aristidoideae

Distribution
from FNA
AZ; CA
[WildflowerSearch map]
[BONAP county map]
Discussion

The range of both varieties of Aristida californica extends from the southwestern United States into northwestern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The subfamily Aristidoideae includes only one tribe, the Aristideae. Other taxonomists have generally included the Aristidoideae, with the Danthonieae and Arundineae, in the Arundinoideae (e.g., Watson et al. 1985; Clayton and Renvoize 1986; Kellogg and Campbell 1987), but Esen and Hilu (1991) demonstrated that Aristida is clearly distinct from the Danthonieae and Arundineae in terms of its prolamins. Subsequent work has provided further support for the monophyly of the three tribes, but their position relative to each other and other members of the PACCAD clade is more equivocal.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline internodes puberulent to nearly lanose
var. californica
1. Cauline internodes glabrous
var. glabrata
Source FNA vol. 25, p. 319. FNA vol. 25, p. 314. Author: Grass Phylogeny Working Group; Kelly W. Allred;.
Parent taxa Poaceae > subfam. Aristidoideae > tribe Aristideae > Aristida Poaceae
Sibling taxa
A. adscensionis, A. arizonica, A. basiramea, A. condensata, A. desmantha, A. dichotoma, A. divaricata, A. floridana, A. gypsophila, A. gyrans, A. havardii, A. lanosa, A. longespica, A. mohrii, A. oligantha, A. palustris, A. pansa, A. patula, A. purpurascens, A. purpurea, A. ramosissima, A. rhizomophora, A. schiedeana, A. simpliciflora, A. spiciformis, A. stricta, A. ternipes, A. tuberculosa
Subordinate taxa
A. californica var. californica, A. californica var. glabrata
Name authority Thurb. Caro
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