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Hill country silverbush

low silverbush

Habit Herbs, perennial, monoecious or dioecious, to 5 dm. Herbs, perennial, usually monoecious, rarely dioecious, to 3 dm.
Stems

erect to ascending, hairy, hairs malpighiaceous.

erect or prostrate, hairy or glabrous, hairs not silvery, malpighiaceous.

Leaves

stipules deciduous, linear-lanceolate, to 1 mm, margins not glandular;

petiole absent;

blade elliptic, 1.5–4 × 0.6–2 cm, margins entire, without glands, surfaces hairy, hairs malpighiaceous.

stipules persistent, ovate to lanceolate, 0.3–1.4 mm, margins not glandular;

petiole 2–10 mm;

blade obovate to linear, 0.5–7 × 0.2–2.2 cm, margins entire, sometimes slightly revolute, without glands, surfaces hairy or glabrous, hairs not silvery, malpighiaceous.

Inflorescences

unisexual, 3–8 cm;

bracts linear-lanceolate, 2–4.5 mm, margins without glands.

bisexual, rarely unisexual (staminate), 0.7–0.9 cm;

bracts ovate to lanceolate, 0.8–2.5 mm, margins without glands.

Flowers

without pink dye when wetted.

without pink dye when wetted.

Staminate flowers

sepals lanceolate to narrowly elliptic, 4–5.5 × 1.1–1.4 mm;

petals cuneate-elliptic to cuneate-obovate, 3.3–4 × 1.6–3 mm, free from androphore;

nectary glands oblong, 0.4–0.5 × 0.3–0.4 mm, adnate to androphore, glabrous;

stamens 10, staminodes 0.

sepals lanceolate, 2.4–3.3 × 0.3–0.8 mm;

petals elliptic, 1.5–2.6 × 0.4–0.8 mm, free from androphore;

nectary glands ovate to linear, 0.4–0.8 × 0.2–0.3 mm, adnate to or free from androphore, glabrous, rarely pubescent;

stamens (7–)10, staminodes 2–3, glabrous.

Pistillate flowers

sepals lanceolate to elliptic, 6–7 × 1.7–3.2 mm, without glands;

petals 0 or 5, linear, 0–1.7 mm;

nectary glands oblong, 0.5 × 0.7 mm, glabrous;

ovary tomentose to lanulose;

styles 1.5–3 mm, tomentose;

stigmas flattened.

sepals elliptic, 4–5.2 × 0.9–2.5 mm, without glands, petals 5, elliptic to filiform, 0.3–1.7 × 0.3–0.4 mm;

nectary glands linear to oblong, 0.3–1.5 × 0.1–0.4 mm, glabrous;

ovary hispidulous, strigillose, or glabrous;

styles 0.6–1.3 mm, glabrous;

stigmas flattened.

Capsules

7–9 mm, tomentose to lanulose.

1.6–4.2 mm, hispidulous, strigillose, or glabrous.

Seeds

3.7–5 mm, smooth, lightly tuberculate, or striate.

1.5–2.5 mm, smooth or rugose-roughened to papillate.

Argythamnia aphoroides

Argythamnia humilis

Phenology Flowering Jan–Jul. Flowering year-round.
Habitat Bluestem-grama grasslands, oak woodlands, calcareous, often rocky, clay or loam soils. Pine-oak forests, desert scrub, grasslands, limestone, sandstone, and clay soils.
Elevation 300–600 m. (1000–2000 ft.) 0–1600 m. (0–5200 ft.)
Distribution
from FNA
TX
[BONAP county map]
from FNA
CO; KS; NM; OK; TX; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas)
[BONAP county map]
Discussion

Argythamnia aphoroides is restricted to the Edwards Plateau.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Glabrous plants have been recognized as Argythamnia laevis (and other combinations of Aphora laevis at different taxonomic ranks). B. L. Turner (2001b) studied this variation and concluded that the glabrous plants do not form distinct populations but instead are individuals in otherwise pubescent populations. Argythamnia humilis var. leiosperma was distinguished by possessing smooth seeds, in contrast to the markedly rugose-roughened seeds of var. humilis. When he described the variety, Waterfall mentioned that var. leiosperma is found in populations of western Oklahoma (Cimarron County) and central Texas (Brown and Hamilton counties). However, this taxon has not been recognized in recent treatments of the plants of Texas (G. M. Diggs et al. 1999; Turner et al. 2003) and Oklahoma (R. J. Tyrl et al. 2010) and is not accepted here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 174. FNA vol. 12, p. 176.
Parent taxa Euphorbiaceae > Argythamnia Euphorbiaceae > Argythamnia
Sibling taxa
A. adenophora, A. argothamnoides, A. argyraea, A. brandegeei, A. claryana, A. cyanophylla, A. humilis, A. lanceolata, A. mercurialina, A. serrata, A. simulans
A. adenophora, A. aphoroides, A. argothamnoides, A. argyraea, A. brandegeei, A. claryana, A. cyanophylla, A. lanceolata, A. mercurialina, A. serrata, A. simulans
Synonyms Ditaxis aphoroides Aphora humilis, A. laevis, A. mercurialina var. pumila, A. humilis var. laevis, A. humilis var. leiosperma, A. laevis, Ditaxis humilis, D. humilis var. leiosperma, D. laevis
Name authority Müller Arg.: Linnaea 34: 146. (1865) — (as Argyrothamnia) (Engelmann & A. Gray) Müller Arg.: Linnaea 34: 147. (1865) — (as Argyrothamnia)
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