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small-leaf false cloak fern

Jones' false cloak-fern, Jones' lipfern

Stem

scales brown.

scales brown to nearly black.

Leaves

7–25 cm.

4–15 cm.

Petiole

brown, 0.75–1.5 mm diam.

dark brown, 0.75–1.5 mm diam.

Blade

deltate to ovate, 3–4-pinnate proximally, leathery, abaxially and adaxially glabrous;

rachis flattened or shallowly grooved adaxially.

ovate-lanceolate, 2–3-pinnate proximally, leathery, abaxially and adaxially glabrous;

rachis rounded to slightly flattened adaxially.

Ultimate segments

articulate, dark color of stalks stopping abruptly at segment bases;

segment margins recurved to revolute, often concealing sporangia;

veins obscure adaxially.

not articulate, dark color of stalks continuing into segment bases abaxially;

segment margins plane to slightly recurved, not concealing sporangia;

veins obscure adaxially.

Sporangia

submarginal, borne on distal 1/3 of secondary veins, containing 64 spores.

submarginal, borne on distal 1/2 of secondary veins, containing 64 spores.

Pinna

costae straight to somewhat flexuous, branches rarely arising from prominent angles.

costae straight or nearly so, branches not arising from prominent angles.

2n

= 54.

= 54, 108.

Argyrochosma microphylla

Argyrochosma jonesii

Phenology Sporulating summer–fall. Sporulating spring–fall.
Habitat Rocky limestone hillsides and cliffs Calcareous cliffs and ledges
Elevation 300–2100 m (1000–6900 ft) 600–1900 m (2000–6200 ft)
Distribution
from FNA
NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT; Mexico in Sonora
[WildflowerSearch map]
[BONAP county map]
Discussion

Argyrochosma microphylla is probably the most distinctive species of Argyrochosma in the flora. Chromosome studies by I. W. Knobloch et al. (1973) suggest that it may include diploid and tetraploid cytotypes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Argyrochosma jonesii includes two sexually reproducing cytotypes. The diploid is known from a few localities in the Sonoran and Mojave Deserts; the tetraploid is found throughout the Mojave Desert and cismontane southern California. Although subtle morphologic differences exist between these cytotypes, they are distinguished primarily by characteristics known to correlate with increases in ploidy level (such as spore size and the length of stomatal guard cells). Further investigation is necessary to determine whether the tetraploid arose through autopolyploidy or hybridization between cryptic species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Argyrochosma Pteridaceae > Argyrochosma
Sibling taxa
A. dealbata, A. fendleri, A. incana, A. jonesii, A. limitanea
A. dealbata, A. fendleri, A. incana, A. limitanea, A. microphylla
Synonyms Pellaea microphylla, Cheilanthes parvifolia, Notholaena parvifolia Notholaena jonesii, Cheilanthes jonesii, Pellaea jonesii
Name authority (Mettenius ex Kuhn) Windham: Amer. Fern J. 77: 40. (1987) (Maxon) Windham: Amer. Fern J. 77: 40. (1987)
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