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spreading sandwort

sabline á feuilles de serpolet, thyme-leaf sandwort

Habit Plants ± strongly perennial, possibly blooming first year, not matted. Plants annual.
Taproots

filiform to moderately thickened;

rhizomes often present, slender, 2–15+ cm.

filiform.

Stems

1–80+, erect or ascending to procumbent or prostrate to trailing, green, 5–60 cm;

internodes terete to angular, 1/3–8+ times as long as leaves, dull, retrorsely pubescent throughout or in lines, hairs minute.

1–100+, erect to ascending or sprawling, green, 3–40+ cm;

internodes terete to ellipsoid, 2–8 times as long as leaves, dull, retrorsely pubescent throughout, sometimes also stipitate-glandular.

Leaves

usually connate basally, with scarious sheath 0.1–0.5 mm, occasionally petiolate (proximal leaves) or sessile;

petiole 2–5 mm;

blade 1-veined, vein prominent abaxially, linear-lanceolate to narrowly elliptic or oblanceolate, 3–35 × 2–14 mm, herbaceous, margins thickened, scarious, shiny, ciliate proximally or throughout, apex obtuse or acute to apiculate, often minutely pustulate, ciliate on margins and adaxial midrib;

axillary leaf clusters absent.

often connate basally, with scarious or mostly herbaceous sheath 0.2–0.3 mm, petiolate (proximal leaves) or usually sessile;

petiole 1–4 mm;

blade 3–5 veined, elliptic to broadly ovate or rarely orbiculate, 2–7 × 1–4 mm, herbaceous, margins flat, herbaceous, dull, ciliate especially proximally, apex acute to acuminate, pustulate, sparsely minutely pubescent or glabrous;

axillary leaf clusters absent.

Inflorescences

axillary, solitary flowers or in proliferating, mostly terminal, leafy, 1–80+-flowered cymes.

terminal, open, leafy, 3–50+-flowered cymes.

Pedicels

erect to ascending (often arcuately so), or straight to widely divergent, often hooked distally in fruit, 2–40 mm, retrorsely pubescent.

erect or ascending in fruit, 1–12 mm, retrorsely pubescent.

Flowers

sepals green, 1–3-veined, 2 lateral veins 1/4–3/4 times as long as midvein, often appearing prominently keeled proximally, lanceolate to ovate (herbaceous portion oblong or lanceolate to ovate), 2–5 mm, to 5.5 mm in fruit, apex acute to acuminate, not pustulate, glabrous;

petals narrowly spatulate to obovate, 1.5–6 mm, 1/2–12/5 times as long as sepals or absent, apex obtuse to rounded, petals sometimes absent.

sepals green, often prominently 3-veined, not keeled, lanceolate to ovate-lanceolate (herbaceous portion narrowly elliptic to broadly lanceolate), 2–3 mm, to 4 mm in fruit, apex narrowly acute to acuminate, ± minutely pustulate, stipitate-glandular;

petals oblong, 0.6–2.7 mm, 1/5–3/4 times as long as sepals, apex obtuse to rounded.

Capsules

± loosely to tightly enclosed by calyx, ovoid, 3–6 mm, 4/5–11/2 times as long as sepals.

loosely enclosed by calyx, ovoid to cylindric-ovoid, 3–3.5 mm, 4/5–11/5 times as long as sepals.

Seeds

8–35, black, suborbicular, slightly compressed, 0.7–0.8 mm, shiny, smooth.

10–15, ashy black, reniform, plump, 0.4–0.6 mm, not shiny, with low-elongate, prominent tubercules.

2n

= 40, 44.

Arenaria lanuginosa

Arenaria serpyllifolia

Distribution
from FNA
AL; AZ; CA; CO; FL; GA; LA; MS; NC; NM; SC; TN; TX; UT; VA; Central America; South America
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[BONAP county map]
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Eurasia; Africa [Introduced, North America; introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4+ (2 in the flora).

Arenaria lanuginosa is morphologically diverse, both in our area and southward into northern South America, and is in serious need of comprehensive study. Other species in subg. Leiosperma (e.g., A. gypsostrata B. L. Turner) that occur in Mexico resemble A. lanuginosa; the nature of those relationships also requires study. We have taken the “conservative approach” of treating the two taxa that occur in the flora area as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3+ (2 in the flora).

Variation in Arenaria serpyllifolia in the broad sense is treated in various ways. The two varieties recognized here have been treated also as subspecies (e.g., A. O. Chater and G. Halliday 1993) or species (e.g., M. N. Abuhadra 2000; B. Jonsell 2001). Jonsell admitted that accepting them as species is questionable; while the morphological differences are slight (see esp. Abuhadra), the ploidy-level difference (2n = 40 in var. serpyllifolia vs. 2n = 20 in var. tenuior) is important.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems often 1-10, prostrate to trailing; inflorescences of solitary, axillary flowers; petals absent or 1/ 3/ 4 times as long as sepals
var. lanuginosa
1. Stems 1-80+, erect or ascending to procumbent; inflorescences of proliferating, leafy, 1-80+-flowered cymes; petals 3/ 2/ 5 times as long as sepals
var. saxosa
1. Capsules ovoid to ovoid-conic, broader at base; seeds 0.5-0.6 mm
var. serpyllifolia
1. Capsules cylindric to cylindric-ovoid, not significantly broader at base (walls nearly straight); seeds ca. 0.4 mm
var. tenuior
Source FNA vol. 5, p. 53. FNA vol. 5, p. 55.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Arenaria Caryophyllaceae > subfam. Alsinoideae > Arenaria
Sibling taxa
A. benthamii, A. humifusa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
A. benthamii, A. humifusa, A. lanuginosa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida
Subordinate taxa
A. lanuginosa var. lanuginosa, A. lanuginosa var. saxosa
A. serpyllifolia var. serpyllifolia, A. serpyllifolia var. tenuior
Synonyms Spergulastrum lanuginosum
Name authority (Michaux) Rohrbach: in C. F. P. von Martius et al., Fl. Bras. 14(2): 274. (1872) Linnaeus: Sp. Pl. 1: 423. (1753)
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