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spreading sandwort

creeping sandwort, low sandwort, sabline rampante

Habit Plants ± strongly perennial, possibly blooming first year, not matted. Plants perennial, matted.
Taproots

filiform to moderately thickened;

rhizomes often present, slender, 2–15+ cm.

Stems

1–80+, erect or ascending to procumbent or prostrate to trailing, green, 5–60 cm;

internodes terete to angular, 1/3–8+ times as long as leaves, dull, retrorsely pubescent throughout or in lines, hairs minute.

5–40+, flowering stems erect to ascending, green, 2–8(–10) cm;

internodes terete to angular, 1/2–4 times as long as leaves, shiny, glabrous or occasionally sparsely stipitate-glandular at distalmost internodes.

Leaves

usually connate basally, with scarious sheath 0.1–0.5 mm, occasionally petiolate (proximal leaves) or sessile;

petiole 2–5 mm;

blade 1-veined, vein prominent abaxially, linear-lanceolate to narrowly elliptic or oblanceolate, 3–35 × 2–14 mm, herbaceous, margins thickened, scarious, shiny, ciliate proximally or throughout, apex obtuse or acute to apiculate, often minutely pustulate, ciliate on margins and adaxial midrib;

axillary leaf clusters absent.

connate basally, with scarious sheath 0.2–0.3 mm, sessile;

blade weakly to prominently 1-veined, narrowly oblong or elliptic to ovate-lanceolate, 2–12 × 1–3 mm, subsucculent, margins thickened, herbaceous, shiny, smooth, apex blunt to acute, prominently pustulate or not, glabrous;

axillary leaf clusters absent.

Inflorescences

axillary, solitary flowers or in proliferating, mostly terminal, leafy, 1–80+-flowered cymes.

terminal, open, leafy, 1–3-flowered cymes or often flowers solitary.

Pedicels

erect to ascending (often arcuately so), or straight to widely divergent, often hooked distally in fruit, 2–40 mm, retrorsely pubescent.

erect in fruit, 1–10 mm, minutely retrorsely pubescent to stipitate-glandular.

Flowers

sepals green, 1–3-veined, 2 lateral veins 1/4–3/4 times as long as midvein, often appearing prominently keeled proximally, lanceolate to ovate (herbaceous portion oblong or lanceolate to ovate), 2–5 mm, to 5.5 mm in fruit, apex acute to acuminate, not pustulate, glabrous;

petals narrowly spatulate to obovate, 1.5–6 mm, 1/2–12/5 times as long as sepals or absent, apex obtuse to rounded, petals sometimes absent.

sepals green, 3-veined, 2 lateral veins 1/2–3/4 times as long as midvein, more prominent in fruit, not keeled, ovate (herbaceous portion ovate), 2.5–4 mm, not enlarging in fruit, apex obtuse or somewhat acute, not pustulate, glabrous;

petals oblanceolate, 3–4 mm, ± 11/2 times as long as sepals, apex acute.

Capsules

± loosely to tightly enclosed by calyx, ovoid, 3–6 mm, 4/5–11/2 times as long as sepals.

tightly enclosed by calyx, broadly ellipsoid, 4–4.5 mm, ca. 11/4 times as long as sepals.

Seeds

8–35, black, suborbicular, slightly compressed, 0.7–0.8 mm, shiny, smooth.

12–18, brown, reniform, compressed, 0.5–0.7 mm, shiny, faintly rugulose.

Tap

-roots filiform;

rhizomes weak, slender, 0.5–15+ cm.

2n

= 40, 44.

= 40, 44 (both Europe).

Arenaria lanuginosa

Arenaria humifusa

Phenology Flowering spring–summer.
Habitat Moist, calcareous gravels and rock crevices
Elevation 0-100 m (0-300 ft)
Distribution
from FNA
AL; AZ; CA; CO; FL; GA; LA; MS; NC; NM; SC; TN; TX; UT; VA; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from FNA
MB; NL; NS; NT; NU; ON; PE; QC; SPM; Greenland; Europe
[BONAP county map]
Discussion

Varieties 4+ (2 in the flora).

Arenaria lanuginosa is morphologically diverse, both in our area and southward into northern South America, and is in serious need of comprehensive study. Other species in subg. Leiosperma (e.g., A. gypsostrata B. L. Turner) that occur in Mexico resemble A. lanuginosa; the nature of those relationships also requires study. We have taken the “conservative approach” of treating the two taxa that occur in the flora area as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arenaria humifusa is similar to A. longipedunculata, but is smaller in some dimensions than that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems often 1-10, prostrate to trailing; inflorescences of solitary, axillary flowers; petals absent or 1/ 3/ 4 times as long as sepals
var. lanuginosa
1. Stems 1-80+, erect or ascending to procumbent; inflorescences of proliferating, leafy, 1-80+-flowered cymes; petals 3/ 2/ 5 times as long as sepals
var. saxosa
Source FNA vol. 5, p. 53. FNA vol. 5, p. 52.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Arenaria Caryophyllaceae > subfam. Alsinoideae > Arenaria
Sibling taxa
A. benthamii, A. humifusa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
A. benthamii, A. lanuginosa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
Subordinate taxa
A. lanuginosa var. lanuginosa, A. lanuginosa var. saxosa
Synonyms Spergulastrum lanuginosum
Name authority (Michaux) Rohrbach: in C. F. P. von Martius et al., Fl. Bras. 14(2): 274. (1872) Wahlenberg: Fl. Lapp., 129. (1812)
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