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spreading sandwort

hilly sandwort

Habit Plants ± strongly perennial, possibly blooming first year, not matted. Plants annual.
Taproots

filiform to moderately thickened;

rhizomes often present, slender, 2–15+ cm.

filiform.

Stems

1–80+, erect or ascending to procumbent or prostrate to trailing, green, 5–60 cm;

internodes terete to angular, 1/3–8+ times as long as leaves, dull, retrorsely pubescent throughout or in lines, hairs minute.

1–40+, erect to ascending, branching, green, 10–30(–50) cm;

internodes ± terete, mostly 1–10 times as long as leaves, dull, retrosely puberulent in 2 lines.

Leaves

usually connate basally, with scarious sheath 0.1–0.5 mm, occasionally petiolate (proximal leaves) or sessile;

petiole 2–5 mm;

blade 1-veined, vein prominent abaxially, linear-lanceolate to narrowly elliptic or oblanceolate, 3–35 × 2–14 mm, herbaceous, margins thickened, scarious, shiny, ciliate proximally or throughout, apex obtuse or acute to apiculate, often minutely pustulate, ciliate on margins and adaxial midrib;

axillary leaf clusters absent.

usually connate proximally, with scarious sheath 0.2–0.4 mm, petiolate (proximal leaves) or sessile;

petiole 2–4 mm;

blade 1-veined, vein ± weak abaxially, ovate (petiolate blades) or elliptic-lanceolate to oblanceolate, 5–15(–20) × 2–4 mm, herbaceous, margins undulate, scarious, shiny, sparsely ciliate especially proximally, apex acute to apiculate or cuspidate, often pustulate, glabrous;

axillary leaf clusters absent.

Inflorescences

axillary, solitary flowers or in proliferating, mostly terminal, leafy, 1–80+-flowered cymes.

terminal, open, leafy, 3–50+-flowered cymes.

Pedicels

erect to ascending (often arcuately so), or straight to widely divergent, often hooked distally in fruit, 2–40 mm, retrorsely pubescent.

erect to reflexed in fruit, 10–40 mm, glabrous.

Flowers

sepals green, 1–3-veined, 2 lateral veins 1/4–3/4 times as long as midvein, often appearing prominently keeled proximally, lanceolate to ovate (herbaceous portion oblong or lanceolate to ovate), 2–5 mm, to 5.5 mm in fruit, apex acute to acuminate, not pustulate, glabrous;

petals narrowly spatulate to obovate, 1.5–6 mm, 1/2–12/5 times as long as sepals or absent, apex obtuse to rounded, petals sometimes absent.

sepals green, 1-veined, vein obscure or prominent proximally, not keeled, elliptic to broadly ovate (herbaceous portion light green, ovate), 2–3.5 mm, to 4.5 mm in fruit, apex acute to acuminate, often pustulate, glabrous;

petals elliptic, 1.5–3 mm, 1/2–3/4 times as long as sepals, apex obtuse.

Capsules

± loosely to tightly enclosed by calyx, ovoid, 3–6 mm, 4/5–11/2 times as long as sepals.

loosely to tightly enclosed by calyx, ovoid, 3–4 mm, 11/5–2 times as long as sepals.

Seeds

8–35, black, suborbicular, slightly compressed, 0.7–0.8 mm, shiny, smooth.

20–30, ashy brown, broadly reniform, compressed, 0.5–0.6 mm, dull, minutely or prominently rounded-tuberculate.

2n

= 40, 44.

Arenaria lanuginosa

Arenaria benthamii

Phenology Flowering late spring.
Habitat Open woodlands, limestone slopes and outcrops
Elevation 0-400 m (0-1300 ft)
Distribution
from FNA
AL; AZ; CA; CO; FL; GA; LA; MS; NC; NM; SC; TN; TX; UT; VA; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; NM; OK; TX; Mexico (Coahuila, Nuevo León)
[BONAP county map]
Discussion

Varieties 4+ (2 in the flora).

Arenaria lanuginosa is morphologically diverse, both in our area and southward into northern South America, and is in serious need of comprehensive study. Other species in subg. Leiosperma (e.g., A. gypsostrata B. L. Turner) that occur in Mexico resemble A. lanuginosa; the nature of those relationships also requires study. We have taken the “conservative approach” of treating the two taxa that occur in the flora area as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems often 1-10, prostrate to trailing; inflorescences of solitary, axillary flowers; petals absent or 1/ 3/ 4 times as long as sepals
var. lanuginosa
1. Stems 1-80+, erect or ascending to procumbent; inflorescences of proliferating, leafy, 1-80+-flowered cymes; petals 3/ 2/ 5 times as long as sepals
var. saxosa
Source FNA vol. 5, p. 53. FNA vol. 5, p. 52.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Arenaria Caryophyllaceae > subfam. Alsinoideae > Arenaria
Sibling taxa
A. benthamii, A. humifusa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
A. humifusa, A. lanuginosa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
Subordinate taxa
A. lanuginosa var. lanuginosa, A. lanuginosa var. saxosa
Synonyms Spergulastrum lanuginosum
Name authority (Michaux) Rohrbach: in C. F. P. von Martius et al., Fl. Bras. 14(2): 274. (1872) Fenzl ex Torrey & A. Gray: Fl. N. Amer. 1: 675. (1840)
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