Arenaria lanuginosa |
Arenaria benthamii |
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spreading sandwort |
hilly sandwort |
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Habit | Plants ± strongly perennial, possibly blooming first year, not matted. | Plants annual. | ||||
Taproots | filiform to moderately thickened; rhizomes often present, slender, 2–15+ cm. |
filiform. |
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Stems | 1–80+, erect or ascending to procumbent or prostrate to trailing, green, 5–60 cm; internodes terete to angular, 1/3–8+ times as long as leaves, dull, retrorsely pubescent throughout or in lines, hairs minute. |
1–40+, erect to ascending, branching, green, 10–30(–50) cm; internodes ± terete, mostly 1–10 times as long as leaves, dull, retrosely puberulent in 2 lines. |
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Leaves | usually connate basally, with scarious sheath 0.1–0.5 mm, occasionally petiolate (proximal leaves) or sessile; petiole 2–5 mm; blade 1-veined, vein prominent abaxially, linear-lanceolate to narrowly elliptic or oblanceolate, 3–35 × 2–14 mm, herbaceous, margins thickened, scarious, shiny, ciliate proximally or throughout, apex obtuse or acute to apiculate, often minutely pustulate, ciliate on margins and adaxial midrib; axillary leaf clusters absent. |
usually connate proximally, with scarious sheath 0.2–0.4 mm, petiolate (proximal leaves) or sessile; petiole 2–4 mm; blade 1-veined, vein ± weak abaxially, ovate (petiolate blades) or elliptic-lanceolate to oblanceolate, 5–15(–20) × 2–4 mm, herbaceous, margins undulate, scarious, shiny, sparsely ciliate especially proximally, apex acute to apiculate or cuspidate, often pustulate, glabrous; axillary leaf clusters absent. |
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Inflorescences | axillary, solitary flowers or in proliferating, mostly terminal, leafy, 1–80+-flowered cymes. |
terminal, open, leafy, 3–50+-flowered cymes. |
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Pedicels | erect to ascending (often arcuately so), or straight to widely divergent, often hooked distally in fruit, 2–40 mm, retrorsely pubescent. |
erect to reflexed in fruit, 10–40 mm, glabrous. |
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Flowers | sepals green, 1–3-veined, 2 lateral veins 1/4–3/4 times as long as midvein, often appearing prominently keeled proximally, lanceolate to ovate (herbaceous portion oblong or lanceolate to ovate), 2–5 mm, to 5.5 mm in fruit, apex acute to acuminate, not pustulate, glabrous; petals narrowly spatulate to obovate, 1.5–6 mm, 1/2–12/5 times as long as sepals or absent, apex obtuse to rounded, petals sometimes absent. |
sepals green, 1-veined, vein obscure or prominent proximally, not keeled, elliptic to broadly ovate (herbaceous portion light green, ovate), 2–3.5 mm, to 4.5 mm in fruit, apex acute to acuminate, often pustulate, glabrous; petals elliptic, 1.5–3 mm, 1/2–3/4 times as long as sepals, apex obtuse. |
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Capsules | ± loosely to tightly enclosed by calyx, ovoid, 3–6 mm, 4/5–11/2 times as long as sepals. |
loosely to tightly enclosed by calyx, ovoid, 3–4 mm, 11/5–2 times as long as sepals. |
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Seeds | 8–35, black, suborbicular, slightly compressed, 0.7–0.8 mm, shiny, smooth. |
20–30, ashy brown, broadly reniform, compressed, 0.5–0.6 mm, dull, minutely or prominently rounded-tuberculate. |
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2n | = 40, 44. |
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Arenaria lanuginosa |
Arenaria benthamii |
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Phenology | Flowering late spring. | |||||
Habitat | Open woodlands, limestone slopes and outcrops | |||||
Elevation | 0-400 m (0-1300 ft) | |||||
Distribution |
AL; AZ; CA; CO; FL; GA; LA; MS; NC; NM; SC; TN; TX; UT; VA; Central America; South America
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LA; NM; OK; TX; Mexico (Coahuila, Nuevo León) |
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Discussion | Varieties 4+ (2 in the flora). Arenaria lanuginosa is morphologically diverse, both in our area and southward into northern South America, and is in serious need of comprehensive study. Other species in subg. Leiosperma (e.g., A. gypsostrata B. L. Turner) that occur in Mexico resemble A. lanuginosa; the nature of those relationships also requires study. We have taken the “conservative approach” of treating the two taxa that occur in the flora area as varieties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 53. | FNA vol. 5, p. 52. | ||||
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Arenaria | Caryophyllaceae > subfam. Alsinoideae > Arenaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Spergulastrum lanuginosum | |||||
Name authority | (Michaux) Rohrbach: in C. F. P. von Martius et al., Fl. Bras. 14(2): 274. (1872) | Fenzl ex Torrey & A. Gray: Fl. N. Amer. 1: 675. (1840) | ||||
Web links |