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spreading sandwort

long-stem sandwort, low sandwort

Habit Plants ± strongly perennial, possibly blooming first year, not matted. Plants perennial, tufted to mat-forming.
Taproots

filiform to moderately thickened;

rhizomes often present, slender, 2–15+ cm.

filiform;

rhizomes slender, 0.5–3 cm.

Stems

1–80+, erect or ascending to procumbent or prostrate to trailing, green, 5–60 cm;

internodes terete to angular, 1/3–8+ times as long as leaves, dull, retrorsely pubescent throughout or in lines, hairs minute.

25–60+, erect to ascending, green, 2–4 cm;

internodes terete, 1–4 times as long as leaves, shiny, minutely glandular-villous.

Leaves

usually connate basally, with scarious sheath 0.1–0.5 mm, occasionally petiolate (proximal leaves) or sessile;

petiole 2–5 mm;

blade 1-veined, vein prominent abaxially, linear-lanceolate to narrowly elliptic or oblanceolate, 3–35 × 2–14 mm, herbaceous, margins thickened, scarious, shiny, ciliate proximally or throughout, apex obtuse or acute to apiculate, often minutely pustulate, ciliate on margins and adaxial midrib;

axillary leaf clusters absent.

briefly connate basally, with herbaceous sheath 0.1–0.3 mm, sessile;

blade obscurely 1-veined, narrowly elliptic to lanceolate or ovate, 2–6 × 1–1.5 mm, subsucculent, margins thickened, herbaceous, shiny, ciliate proximally, apex acute to apiculate, pustulate, glabrous;

axillary leaf clusters absent.

Inflorescences

axillary, solitary flowers or in proliferating, mostly terminal, leafy, 1–80+-flowered cymes.

terminal, solitary flowers.

Pedicels

erect to ascending (often arcuately so), or straight to widely divergent, often hooked distally in fruit, 2–40 mm, retrorsely pubescent.

erect in fruit, 10–20 mm, densely glandular-villous.

Flowers

sepals green, 1–3-veined, 2 lateral veins 1/4–3/4 times as long as midvein, often appearing prominently keeled proximally, lanceolate to ovate (herbaceous portion oblong or lanceolate to ovate), 2–5 mm, to 5.5 mm in fruit, apex acute to acuminate, not pustulate, glabrous;

petals narrowly spatulate to obovate, 1.5–6 mm, 1/2–12/5 times as long as sepals or absent, apex obtuse to rounded, petals sometimes absent.

sepals green, obscurely veined, not keeled, ovate (herbaceous portion ovate), 3.5–4 mm, not enlarging in fruit, apex acute to acuminate, not pustulate, minutely stipitate-glandular proximally;

petals broadly elliptic, 2.5–3.5 mm, 11/4–11/2 times as long as sepals, apex rounded.

Capsules

± loosely to tightly enclosed by calyx, ovoid, 3–6 mm, 4/5–11/2 times as long as sepals.

tightly enclosed by calyx, ellipsoid, 4–5 mm, 1–11/5 times as long as sepals.

Seeds

8–35, black, suborbicular, slightly compressed, 0.7–0.8 mm, shiny, smooth.

13–20, brown, reniform, compressed, 0.7–0.9 mm, shiny, rugulose (30x).

2n

= 40, 44.

= 40, 80.

Arenaria lanuginosa

Arenaria longipedunculata

Phenology Flowering spring–summer.
Habitat Gravelly, moist, montane areas, open alpine woods
Elevation 50-1000 m (200-3300 ft)
Distribution
from FNA
AL; AZ; CA; CO; FL; GA; LA; MS; NC; NM; SC; TN; TX; UT; VA; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; NT; YT; Asia
[BONAP county map]
Discussion

Varieties 4+ (2 in the flora).

Arenaria lanuginosa is morphologically diverse, both in our area and southward into northern South America, and is in serious need of comprehensive study. Other species in subg. Leiosperma (e.g., A. gypsostrata B. L. Turner) that occur in Mexico resemble A. lanuginosa; the nature of those relationships also requires study. We have taken the “conservative approach” of treating the two taxa that occur in the flora area as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arenaria longipedunculata was for many years included in A. humifusa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems often 1-10, prostrate to trailing; inflorescences of solitary, axillary flowers; petals absent or 1/ 3/ 4 times as long as sepals
var. lanuginosa
1. Stems 1-80+, erect or ascending to procumbent; inflorescences of proliferating, leafy, 1-80+-flowered cymes; petals 3/ 2/ 5 times as long as sepals
var. saxosa
Source FNA vol. 5, p. 53. FNA vol. 5, p. 54.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Arenaria Caryophyllaceae > subfam. Alsinoideae > Arenaria
Sibling taxa
A. benthamii, A. humifusa, A. livermorensis, A. longipedunculata, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
A. benthamii, A. humifusa, A. lanuginosa, A. livermorensis, A. ludens, A. paludicola, A. pseudofrigida, A. serpyllifolia
Subordinate taxa
A. lanuginosa var. lanuginosa, A. lanuginosa var. saxosa
Synonyms Spergulastrum lanuginosum
Name authority (Michaux) Rohrbach: in C. F. P. von Martius et al., Fl. Bras. 14(2): 274. (1872) Hultén: Bot. Not. 119: 313, fig. 1. (1966)
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