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green-leaf manzanita

Eastwood manzanita, Eastwood's manzanita

Habit Shrubs, erect or mound-forming, 1–3 m; burl usually absent, sometimes flat, obscure; twigs usually densely short-hairy with golden glands on tips of hairs, rarely short white-hairy and eglandular. Shrubs, erect or mound-forming, 1–3 m; burl present; twigs tomentose to short-hairy, with eglandular hairs or with long, clear, pink or dark, glandular hairs.
Leaves

petiole 7–15 mm;

blade bright green (lightly gray-green if short-hairy), shiny, widely ovate to orbiculate, 2.5–6 × 1.5–4 cm, base rounded, truncate, or slightly lobed, (not clasping), margins entire, plane, surfaces smooth, glabrous or, rarely, short-hairy.

petiole 5–10 mm;

blade bright green, slightly glaucous to strongly gray-glaucous, shiny or dull, elliptic to ovate, 2–4.5 × 1–2.5 cm, base cuneate to rounded, sometimes ± lobed or truncate, margins entire or toothed, plane, surfaces papillate, scabrous or smooth, glabrous or glandular-puberulent to -hairy, puberulent, or finely tomentose, sometimes glabrescent; (abaxial surface sometimes with more stomata than adaxial surface).

Inflorescences

panicles, 2–8-branched;

immature inflorescence pendent, branches spreading, axis 1.5–3 cm, 1+ mm diam., hairy with golden glands on tips of hairs or short-hairy and eglandular;

bracts appressed with incurved tips, scalelike, deltate, 4–6 mm, apex acuminate, surfaces usually densely tomentose with golden glands on tips of hairs, rarely short white-hairy and eglandular.

panicles, 3–6-branched;

immature inflorescence pendent, (branches crowded, not concealed by bracts), axis 1–3 cm, 1+ mm diam., short-hairy to hairy, with or without glandular hairs;

bracts not appressed, variable, from scalelike and awl-like to leaflike and deltate, lanceolate, or broadly lanceolate (at least at base), 3–10 mm, apex acute, surfaces hairy or finely glandular-hairy.

Pedicels

2–7 mm, glabrous or white-hairy.

3–10 mm, sparsely hairy to finely glandular-hairy.

Flowers

corolla mostly pink, conic to urceolate;

ovary glabrous or white-hairy.

corolla white to pink, conic to urceolate;

ovary densely white-hairy, sometimes glandular.

Fruits

depressed-globose, sometimes subglobose, 7–10 mm diam., glabrous.

slightly (or markedly) depressed-globose, 6–10 mm diam., glabrous or finely glandular-hairy, (viscid).

Stones

distinct.

usually mostly distinct, sometimes connate.

2n

= 26.

Arctostaphylos patula

Arctostaphylos glandulosa

Phenology Flowering late spring–early summer.
Habitat Mountain chaparral and forests
Elevation 400-3000 m (1300-9800 ft)
Distribution
from FNA
AZ; CA; CO; MT; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Arctostaphylos patula is abundant and widespread in western North America as a dominant in montane chaparral, pine forest gaps, and high-elevation arid-steppe and canyon-land environments. Populations throughout western North America are characterized by twigs and inflorescence parts covered with relatively short hairs tipped with golden glands. In the central to northern Sierra Nevada, mixed with the widespread form are individuals that are eglandular and have a cover of relatively short, whitish hairs on the stems and inflorescences. Similarly, throughout most of its range, A. patula is nonsprouting after fire, and in areas characterized by winter snow cover it layers and creates broad, low mounds. In much of California, it typically sprouts after fires from obscure and flattened burls, forming circles of erect sprouts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 10 (8 in the flora).

Arctostaphylos glandulosa occurs in the California Floristic Province from southern Oregon to northern Baja California within the Coast, Transverse, and Peninsular ranges.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Twigs, bracts, and/or immature inflorescence axes glandular-hairy
→ 2
1. Twigs, bracts, and immature inflorescence axes not glandular-hairy
→ 4
2. Leaf blades strongly white-glaucous.
subsp. leucophylla
2. Leaf blades green to gray-green, not glaucous to moderately glaucous
→ 3
3. Twigs glandular-hairy; immature inflorescences usually glandular-hairy.
subsp. glandulosa
3. Twigs not glandular-hairy; immature inflorescences glandular-hairy; (interior and eastern side of Santa Lucia Mountains).
subsp. howellii
4. Twigs short-hairy and with longer, white hairs.
subsp. mollis
4. Twigs short-hairy, not glandular-hairy
→ 5
5. Leaf blades strongly white-glaucous.
subsp. adamsii
5. Leaf blades dark green to moderately gray-green
→ 6
6. Stones mostly connate into single or weakly connate sphere.
subsp. gabrielensis
6. Stones usually distinct
→ 7
7. Leaf blades dark green (margins often reddish); fruits markedly flattened into depressed-globose shape.
subsp. crassifolia
7. Leaf blades green or gray-green; fruits only slightly flattened into depressed-globose shape.
subsp. cushingiana
Source FNA vol. 8, p. 416. FNA vol. 8, p. 442.
Parent taxa Ericaceae > subfam. Arbutoideae > Arctostaphylos Ericaceae > subfam. Arbutoideae > Arctostaphylos
Sibling taxa
A. andersonii, A. auriculata, A. bakeri, A. canescens, A. catalinae, A. columbiana, A. confertiflora, A. crustacea, A. cruzensis, A. densiflora, A. edmundsii, A. franciscana, A. gabilanensis, A. glandulosa, A. glauca, A. glutinosa, A. hispidula, A. hookeri, A. hooveri, A. imbricata, A. insularis, A. klamathensis, A. luciana, A. malloryi, A. manzanita, A. mewukka, A. montana, A. montaraensis, A. montereyensis, A. morroensis, A. myrtifolia, A. nevadensis, A. nissenana, A. nortensis, A. nummularia, A. obispoensis, A. ohloneana, A. osoensis, A. otayensis, A. pacifica, A. pajaroensis, A. pallida, A. parryana, A. pechoensis, A. pilosula, A. pringlei, A. pumila, A. pungens, A. purissima, A. rainbowensis, A. refugioensis, A. regismontana, A. rudis, A. sensitiva, A. silvicola, A. stanfordiana, A. tomentosa, A. uva-ursi, A. virgata, A. viridissima, A. viscida
A. andersonii, A. auriculata, A. bakeri, A. canescens, A. catalinae, A. columbiana, A. confertiflora, A. crustacea, A. cruzensis, A. densiflora, A. edmundsii, A. franciscana, A. gabilanensis, A. glauca, A. glutinosa, A. hispidula, A. hookeri, A. hooveri, A. imbricata, A. insularis, A. klamathensis, A. luciana, A. malloryi, A. manzanita, A. mewukka, A. montana, A. montaraensis, A. montereyensis, A. morroensis, A. myrtifolia, A. nevadensis, A. nissenana, A. nortensis, A. nummularia, A. obispoensis, A. ohloneana, A. osoensis, A. otayensis, A. pacifica, A. pajaroensis, A. pallida, A. parryana, A. patula, A. pechoensis, A. pilosula, A. pringlei, A. pumila, A. pungens, A. purissima, A. rainbowensis, A. refugioensis, A. regismontana, A. rudis, A. sensitiva, A. silvicola, A. stanfordiana, A. tomentosa, A. uva-ursi, A. virgata, A. viridissima, A. viscida
Subordinate taxa
A. glandulosa subsp. adamsii, A. glandulosa subsp. crassifolia, A. glandulosa subsp. cushingiana, A. glandulosa subsp. gabrielensis, A. glandulosa subsp. glandulosa, A. glandulosa subsp. howellii, A. glandulosa subsp. leucophylla, A. glandulosa subsp. mollis
Synonyms A. acutifolia, A. parryana var. pinetorum, A. patula var. coalescens, A. patula subsp. platyphylla, A. platyphylla
Name authority Greene: Pittonia 2: 171. 1891 , Eastwood: Proc. Calif. Acad. Sci., ser. 3, 1: 82. 1897 ,
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