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green-leaf manzanita

bristly manzanita, Columbia manzanita, hairy manzanita, redwood manzanita

Habit Shrubs, erect or mound-forming, 1–3 m; burl usually absent, sometimes flat, obscure; twigs usually densely short-hairy with golden glands on tips of hairs, rarely short white-hairy and eglandular. Shrubs or trees, erect, 2–5 m; burl absent; twigs densely short-hairy, usually with long, white hairs, often glandular.
Leaves

petiole 7–15 mm;

blade bright green (lightly gray-green if short-hairy), shiny, widely ovate to orbiculate, 2.5–6 × 1.5–4 cm, base rounded, truncate, or slightly lobed, (not clasping), margins entire, plane, surfaces smooth, glabrous or, rarely, short-hairy.

petiole 4–10 mm;

blade dark green, dull, lanceolate-ovate to narrowly oblong-ovate, 4–6 × 2–3 cm, base cuneate to ± rounded, margins entire, plane, surfaces ± papillate, finely scabrous, finely hairy, sparsely glandular-hairy.

Inflorescences

panicles, 2–8-branched;

immature inflorescence pendent, branches spreading, axis 1.5–3 cm, 1+ mm diam., hairy with golden glands on tips of hairs or short-hairy and eglandular;

bracts appressed with incurved tips, scalelike, deltate, 4–6 mm, apex acuminate, surfaces usually densely tomentose with golden glands on tips of hairs, rarely short white-hairy and eglandular.

panicles, 3–8-branched;

immature inflorescence pendent, branches ± spreading, axis 1.5–2.5 cm, 1+ mm diam., densely short-hairy, usually with long, white hairs, sometimes glandular;

bracts not appressed, (green), leaflike, oblong-lanceolate, 10–18 mm, apex acuminate, surfaces finely glandular-hairy.

Pedicels

2–7 mm, glabrous or white-hairy.

2–4 mm, finely glandular-hairy.

Flowers

corolla mostly pink, conic to urceolate;

ovary glabrous or white-hairy.

corolla white, conic to urceolate;

ovary densely white-hairy, sometimes sparsely glandular.

Fruits

depressed-globose, sometimes subglobose, 7–10 mm diam., glabrous.

depressed-globose, 8–11 mm diam., sparsely hairy.

Stones

distinct.

distinct.

2n

= 26.

= 26.

Arctostaphylos patula

Arctostaphylos columbiana

Phenology Flowering late spring–early summer. Flowering winter–early spring.
Habitat Mountain chaparral and forests Chaparral, gaps and margins of conifer forests along coast, sometimes extending inland, open areas around rock outcrops
Elevation 400-3000 m (1300-9800 ft) 0-1000(-1400) m (0-3300(-4600) ft)
Distribution
from FNA
AZ; CA; CO; MT; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Arctostaphylos patula is abundant and widespread in western North America as a dominant in montane chaparral, pine forest gaps, and high-elevation arid-steppe and canyon-land environments. Populations throughout western North America are characterized by twigs and inflorescence parts covered with relatively short hairs tipped with golden glands. In the central to northern Sierra Nevada, mixed with the widespread form are individuals that are eglandular and have a cover of relatively short, whitish hairs on the stems and inflorescences. Similarly, throughout most of its range, A. patula is nonsprouting after fire, and in areas characterized by winter snow cover it layers and creates broad, low mounds. In much of California, it typically sprouts after fires from obscure and flattened burls, forming circles of erect sprouts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arctostaphylos columbiana is widespread near the coast from northern California to southern British Columbia; it extends inland along the Columbia River Gorge in Oregon and Washington, and inland in Oregon to the western base of the Cascades. Some variation in twig indument occurs in the prominence of longer, stiff hairs, and in the degree of glandulosity. Some plants along the immediate coast of northern California into Oregon lack the longer hairs and have been distinguished as var. tracyi. Hybrids with A. uva-ursi are low shrubs (0.5–1 m) with intermediate vegetative characters. Referred to as A. ×media Piper, these hybrids have been reported from British Columbia, California, Oregon, and Washington.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 416. FNA vol. 8, p. 425.
Parent taxa Ericaceae > subfam. Arbutoideae > Arctostaphylos Ericaceae > subfam. Arbutoideae > Arctostaphylos
Sibling taxa
A. andersonii, A. auriculata, A. bakeri, A. canescens, A. catalinae, A. columbiana, A. confertiflora, A. crustacea, A. cruzensis, A. densiflora, A. edmundsii, A. franciscana, A. gabilanensis, A. glandulosa, A. glauca, A. glutinosa, A. hispidula, A. hookeri, A. hooveri, A. imbricata, A. insularis, A. klamathensis, A. luciana, A. malloryi, A. manzanita, A. mewukka, A. montana, A. montaraensis, A. montereyensis, A. morroensis, A. myrtifolia, A. nevadensis, A. nissenana, A. nortensis, A. nummularia, A. obispoensis, A. ohloneana, A. osoensis, A. otayensis, A. pacifica, A. pajaroensis, A. pallida, A. parryana, A. pechoensis, A. pilosula, A. pringlei, A. pumila, A. pungens, A. purissima, A. rainbowensis, A. refugioensis, A. regismontana, A. rudis, A. sensitiva, A. silvicola, A. stanfordiana, A. tomentosa, A. uva-ursi, A. virgata, A. viridissima, A. viscida
A. andersonii, A. auriculata, A. bakeri, A. canescens, A. catalinae, A. confertiflora, A. crustacea, A. cruzensis, A. densiflora, A. edmundsii, A. franciscana, A. gabilanensis, A. glandulosa, A. glauca, A. glutinosa, A. hispidula, A. hookeri, A. hooveri, A. imbricata, A. insularis, A. klamathensis, A. luciana, A. malloryi, A. manzanita, A. mewukka, A. montana, A. montaraensis, A. montereyensis, A. morroensis, A. myrtifolia, A. nevadensis, A. nissenana, A. nortensis, A. nummularia, A. obispoensis, A. ohloneana, A. osoensis, A. otayensis, A. pacifica, A. pajaroensis, A. pallida, A. parryana, A. patula, A. pechoensis, A. pilosula, A. pringlei, A. pumila, A. pungens, A. purissima, A. rainbowensis, A. refugioensis, A. regismontana, A. rudis, A. sensitiva, A. silvicola, A. stanfordiana, A. tomentosa, A. uva-ursi, A. virgata, A. viridissima, A. viscida
Synonyms A. acutifolia, A. parryana var. pinetorum, A. patula var. coalescens, A. patula subsp. platyphylla, A. platyphylla A. columbiana var. tracyi, A. tracyi
Name authority Greene: Pittonia 2: 171. 1891 , Piper: in C. V. Piper and R. K. Beattie, Fl. N.W. Coast, 279. 1915 ,
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