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Eastwood manzanita, Eastwood's manzanita

Photo is of parent taxon

Cushing manzanita, Cushing's manzanita

Habit Shrubs, erect or mound-forming, 1–3 m; burl present; twigs tomentose to short-hairy, with eglandular hairs or with long, clear, pink or dark, glandular hairs.
Twigs

sparsely short-hairy, not glandular-hairy.

Leaves

petiole 5–10 mm;

blade bright green, slightly glaucous to strongly gray-glaucous, shiny or dull, elliptic to ovate, 2–4.5 × 1–2.5 cm, base cuneate to rounded, sometimes ± lobed or truncate, margins entire or toothed, plane, surfaces papillate, scabrous or smooth, glabrous or glandular-puberulent to -hairy, puberulent, or finely tomentose, sometimes glabrescent; (abaxial surface sometimes with more stomata than adaxial surface).

blades green or gray-green, base cuneate to rounded, surfaces smooth, glabrous or sparsely puberulent.

Inflorescences

panicles, 3–6-branched;

immature inflorescence pendent, (branches crowded, not concealed by bracts), axis 1–3 cm, 1+ mm diam., short-hairy to hairy, with or without glandular hairs;

bracts not appressed, variable, from scalelike and awl-like to leaflike and deltate, lanceolate, or broadly lanceolate (at least at base), 3–10 mm, apex acute, surfaces hairy or finely glandular-hairy.

immature inflorescence axis densely white-hairy;

bracts densely white-hairy.

Pedicels

3–10 mm, sparsely hairy to finely glandular-hairy.

densely white-hairy.

Flowers

corolla white to pink, conic to urceolate;

ovary densely white-hairy, sometimes glandular.

Ovaries

densely white-hairy.

Fruits

slightly (or markedly) depressed-globose, 6–10 mm diam., glabrous or finely glandular-hairy, (viscid).

slightly flattened depressed-globose.

Stones

usually mostly distinct, sometimes connate.

usually distinct.

Arctostaphylos glandulosa

Arctostaphylos glandulosa subsp. cushingiana

Phenology Flowering winter–early spring.
Habitat Pure and mixed chaparral
Elevation 300-1900 m (1000-6200 ft)
Distribution
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[BONAP county map]
Discussion

Subspecies 10 (8 in the flora).

Arctostaphylos glandulosa occurs in the California Floristic Province from southern Oregon to northern Baja California within the Coast, Transverse, and Peninsular ranges.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies cushingiana is widespread in mixed populations with subsp. glandulosa from Humboldt County in the North Coast Range south to the Peninsular Range, and extending into northern Baja California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Twigs, bracts, and/or immature inflorescence axes glandular-hairy
→ 2
1. Twigs, bracts, and immature inflorescence axes not glandular-hairy
→ 4
2. Leaf blades strongly white-glaucous.
subsp. leucophylla
2. Leaf blades green to gray-green, not glaucous to moderately glaucous
→ 3
3. Twigs glandular-hairy; immature inflorescences usually glandular-hairy.
subsp. glandulosa
3. Twigs not glandular-hairy; immature inflorescences glandular-hairy; (interior and eastern side of Santa Lucia Mountains).
subsp. howellii
4. Twigs short-hairy and with longer, white hairs.
subsp. mollis
4. Twigs short-hairy, not glandular-hairy
→ 5
5. Leaf blades strongly white-glaucous.
subsp. adamsii
5. Leaf blades dark green to moderately gray-green
→ 6
6. Stones mostly connate into single or weakly connate sphere.
subsp. gabrielensis
6. Stones usually distinct
→ 7
7. Leaf blades dark green (margins often reddish); fruits markedly flattened into depressed-globose shape.
subsp. crassifolia
7. Leaf blades green or gray-green; fruits only slightly flattened into depressed-globose shape.
subsp. cushingiana
Source FNA vol. 8, p. 442. FNA vol. 8, p. 444.
Parent taxa Ericaceae > subfam. Arbutoideae > Arctostaphylos Ericaceae > subfam. Arbutoideae > Arctostaphylos > Arctostaphylos glandulosa
Sibling taxa
A. andersonii, A. auriculata, A. bakeri, A. canescens, A. catalinae, A. columbiana, A. confertiflora, A. crustacea, A. cruzensis, A. densiflora, A. edmundsii, A. franciscana, A. gabilanensis, A. glauca, A. glutinosa, A. hispidula, A. hookeri, A. hooveri, A. imbricata, A. insularis, A. klamathensis, A. luciana, A. malloryi, A. manzanita, A. mewukka, A. montana, A. montaraensis, A. montereyensis, A. morroensis, A. myrtifolia, A. nevadensis, A. nissenana, A. nortensis, A. nummularia, A. obispoensis, A. ohloneana, A. osoensis, A. otayensis, A. pacifica, A. pajaroensis, A. pallida, A. parryana, A. patula, A. pechoensis, A. pilosula, A. pringlei, A. pumila, A. pungens, A. purissima, A. rainbowensis, A. refugioensis, A. regismontana, A. rudis, A. sensitiva, A. silvicola, A. stanfordiana, A. tomentosa, A. uva-ursi, A. virgata, A. viridissima, A. viscida
A. glandulosa subsp. adamsii, A. glandulosa subsp. crassifolia, A. glandulosa subsp. gabrielensis, A. glandulosa subsp. glandulosa, A. glandulosa subsp. howellii, A. glandulosa subsp. leucophylla, A. glandulosa subsp. mollis
Subordinate taxa
A. glandulosa subsp. adamsii, A. glandulosa subsp. crassifolia, A. glandulosa subsp. cushingiana, A. glandulosa subsp. gabrielensis, A. glandulosa subsp. glandulosa, A. glandulosa subsp. howellii, A. glandulosa subsp. leucophylla, A. glandulosa subsp. mollis
Synonyms A. cushingiana, A. glandulosa var. cushingiana
Name authority Eastwood: Proc. Calif. Acad. Sci., ser. 3, 1: 82. 1897 , (Eastwood) J. E. Keeley: Madroño 54: 59. (2007)
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