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Photo is of parent taxon

blushing rock-cress, hairy fruit rock cress, hairy rockcress

blushing rock-cress, cream-flower rock-cress, hairy eared-rockcress, hairy rockcress, slender rock cress

Habit Biennials or perennials; (caudex branched); usually densely hirsute (at least basally), rarely glabrescent, trichomes simple mixed with stalked or sessile, forked ones.
Stems

usually hirsute basally, rarely glabrescent, trichomes often simple, sometimes branched.

simple or several from base (rosette), erect, often branched distally, 1–8 dm, (pilose with trichomes appressed, malpighiaceous, or minutely stalked, forked, or hirsute basally with trichomes simple and minutely stalked, forked, sometimes almost exclusively pubescent with forked submalpighiaceous trichomes).

Basal leaves

blade surfaces sparsely to densely pubescent, trichomes simple and stalked.

petiole 0.5–2 cm, (ciliate or not);

blade spatulate, oblanceolate, or oblong, (0.8–)1.5–8 cm × (5–)10–25 mm, margins entire, repand, or dentate, apex obtuse or acute, surfaces sparsely to densely pubescent, trichomes sessile or stalked, simple or forked, and/or stellate.

Cauline leaves

(7–)10–45(–61), (overlapping or not);

blade ovate to oblong or lanceolate, rarely linear, (1–)1.5–6(–8) cm × (1–)3–20(–25) mm, base subcordate or auriculate (auricles obtuse or subacute), margins dentate or entire, apex acute or obtuse, surfaces hirsute or adaxially glabrescent.

Racemes

often simple.

Flowers

sepals oblong, 2.5–4 × 0.5–1.5 mm, lateral pair not saccate basally;

petals white, linear-oblanceolate or narrowly spatulate, 3.5–5(–5.5) × 1–2(–2.5) mm, apex obtuse;

filaments 2.5–4 mm;

anthers oblong, 0.7–1 mm.

Fruiting pedicels

erect to erect-ascending, (2–)3–8(–12) mm (glabrous or sparsely pubescent).

Fruits

(3.5–)4–5.8(–6) cm;

style (0.2–)0.5–1 mm, (rarely stout).

erect to erect-ascending, (often appressed to rachis), torulose, (3.5–)4–6(–6.5) cm × 0.8–1(–1.2) mm;

valves each with obscure or somewhat prominent midvein extending to the middle;

ovules (54–)60–86 per ovary;

style (0.2–)0.5–1(–1.3) mm, (slender).

Seeds

narrowly winged throughout, oblong or suborbicular, (0.8–)1–1.5(–1.7) × 0.8–1.3 mm;

wing to 0.2 mm wide distally.

2n

= 32.

Arabis pycnocarpa var. pycnocarpa

Arabis pycnocarpa

Phenology Flowering Mar–Jul.
Habitat Bluffs, cliffs, ledges, rocky hillsides, open woods, bottom lands, gravel bars, meadows, streamsides, upland prairies, grassy swales, hillsides, stream bottoms
Elevation 0-2500 m (0-8200 ft)
Distribution
from FNA
AK; AZ; CA; CO; CT; IA; ID; KS; MA; ME; MI; MN; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; QC; SK; YT; Asia (China, Japan, Russian Far East)
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; TN; UT; VA; VT; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; QC; SK; YT; Asia
[WildflowerSearch map]
[BONAP county map]
Discussion

Some populations in northern Indiana and northern Illinois, [e.g., Herman 8790 (Jo Davies County) and Friesner 19072 (Elkhart County), both at GH] most likely represent hybrids between the two varieties of Arabis pycnocarpa. Trichomes borne proximally on stems of these plants are a mixture of forked and simple. It is expected that hybrids can be found where the ranges of the two taxa overlap.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

M. Hopkins (1937) synthesized earlier works on Arabis pycnocarpa and concluded that it is different from the European A. hirsuta. Both R. C. Rollins (1941, 1993) and G. A. Mulligan (1996) considered the North American and European plants different varieties of A. hirsuta; Mulligan recognized var. hirsuta in North America and Rollins did not. After examining thousands of specimens from Europe, Asia, and North America, I conclude that Hopkins was correct in treating the North American plants as a different species, A. pycnocarpa. The European A. hirsuta is a diploid (2n = 16) that has fruits 1.5–4 cm, stout styles 0.1–0.5 mm, prominent midvein extending the full length of the fruit valve, and 30–40(–44) ovules/seeds per ovary/fruit. By contrast, A. pycnocarpa is a tetraploid (2n = 32) that has fruits (3.5–)4–6(–6.5) cm, often slender styles (0.2–)0.5–1(–1.3) mm, obscure midvein hardly extending to the middle of the fruit valve, and ovules/seeds (54–)60–86 per ovary/fruit. Hopkins listed other differences in the number of cauline leaves and the development of seed wing, but these do not hold. Mulligan treated the perennial North American plants with minute styles as var. hirsuta and the biennial ones with longer styles as var. pycnocarpa. Habit and style length are not correlated at all, and one finds both biennials and perennials flowering in the same population.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually hirsute basally, rarely glabrescent, trichomes often simple, sometimes branched; basal leaf blades: surfaces with simple and stalked trichomes.
var. pycnocarpa
1. Stems pilose, trichomes appressed, malpighiaceous, or minutely stalked, forked; basal leaf blades: surfaces with sessile, forked, and/or stellatetrichomes.
var. adpressipilis
Source FNA vol. 7, p. 260. FNA vol. 7, p. 259.
Parent taxa Brassicaceae > tribe Arabideae > Arabis > Arabis pycnocarpa Brassicaceae > tribe Arabideae > Arabis
Sibling taxa
A. pycnocarpa var. adpressipilis
A. aculeolata, A. alpina, A. blepharophylla, A. caucasica, A. crucisetosa, A. eschscholtziana, A. furcata, A. georgiana, A. mcdonaldiana, A. modesta, A. nuttallii, A. olympica, A. oregana, A. patens
Subordinate taxa
A. pycnocarpa var. adpressipilis, A. pycnocarpa var. pycnocarpa
Synonyms A. hirsuta var. minshallii, A. pycnocarpa var. reducta A. hirsuta subsp. pycnocarpa, A. hirsuta var. pycnocarpa
Name authority unknown M. Hopkins: Rhodora 39: 112, plate 458, figs. 1–3. (1937)
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