Arabis pycnocarpa var. pycnocarpa |
Arabis pycnocarpa |
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blushing rock-cress, hairy fruit rock cress, hairy rockcress |
blushing rock-cress, cream-flower rock-cress, hairy eared-rockcress, hairy rockcress, slender rock cress |
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Habit | Biennials or perennials; (caudex branched); usually densely hirsute (at least basally), rarely glabrescent, trichomes simple mixed with stalked or sessile, forked ones. | |||||
Stems | usually hirsute basally, rarely glabrescent, trichomes often simple, sometimes branched. |
simple or several from base (rosette), erect, often branched distally, 1–8 dm, (pilose with trichomes appressed, malpighiaceous, or minutely stalked, forked, or hirsute basally with trichomes simple and minutely stalked, forked, sometimes almost exclusively pubescent with forked submalpighiaceous trichomes). |
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Basal leaves | blade surfaces sparsely to densely pubescent, trichomes simple and stalked. |
petiole 0.5–2 cm, (ciliate or not); blade spatulate, oblanceolate, or oblong, (0.8–)1.5–8 cm × (5–)10–25 mm, margins entire, repand, or dentate, apex obtuse or acute, surfaces sparsely to densely pubescent, trichomes sessile or stalked, simple or forked, and/or stellate. |
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Cauline leaves | (7–)10–45(–61), (overlapping or not); blade ovate to oblong or lanceolate, rarely linear, (1–)1.5–6(–8) cm × (1–)3–20(–25) mm, base subcordate or auriculate (auricles obtuse or subacute), margins dentate or entire, apex acute or obtuse, surfaces hirsute or adaxially glabrescent. |
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Racemes | often simple. |
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Flowers | sepals oblong, 2.5–4 × 0.5–1.5 mm, lateral pair not saccate basally; petals white, linear-oblanceolate or narrowly spatulate, 3.5–5(–5.5) × 1–2(–2.5) mm, apex obtuse; filaments 2.5–4 mm; anthers oblong, 0.7–1 mm. |
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Fruiting pedicels | erect to erect-ascending, (2–)3–8(–12) mm (glabrous or sparsely pubescent). |
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Fruits | (3.5–)4–5.8(–6) cm; style (0.2–)0.5–1 mm, (rarely stout). |
erect to erect-ascending, (often appressed to rachis), torulose, (3.5–)4–6(–6.5) cm × 0.8–1(–1.2) mm; valves each with obscure or somewhat prominent midvein extending to the middle; ovules (54–)60–86 per ovary; style (0.2–)0.5–1(–1.3) mm, (slender). |
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Seeds | narrowly winged throughout, oblong or suborbicular, (0.8–)1–1.5(–1.7) × 0.8–1.3 mm; wing to 0.2 mm wide distally. |
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2n | = 32. |
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Arabis pycnocarpa var. pycnocarpa |
Arabis pycnocarpa |
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Phenology | Flowering Mar–Jul. | |||||
Habitat | Bluffs, cliffs, ledges, rocky hillsides, open woods, bottom lands, gravel bars, meadows, streamsides, upland prairies, grassy swales, hillsides, stream bottoms | |||||
Elevation | 0-2500 m (0-8200 ft) | |||||
Distribution |
AK; AZ; CA; CO; CT; IA; ID; KS; MA; ME; MI; MN; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; QC; SK; YT; Asia (China, Japan, Russian Far East) |
AK; AZ; CA; CO; CT; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; TN; UT; VA; VT; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; QC; SK; YT; Asia
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Discussion | Some populations in northern Indiana and northern Illinois, [e.g., Herman 8790 (Jo Davies County) and Friesner 19072 (Elkhart County), both at GH] most likely represent hybrids between the two varieties of Arabis pycnocarpa. Trichomes borne proximally on stems of these plants are a mixture of forked and simple. It is expected that hybrids can be found where the ranges of the two taxa overlap. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). M. Hopkins (1937) synthesized earlier works on Arabis pycnocarpa and concluded that it is different from the European A. hirsuta. Both R. C. Rollins (1941, 1993) and G. A. Mulligan (1996) considered the North American and European plants different varieties of A. hirsuta; Mulligan recognized var. hirsuta in North America and Rollins did not. After examining thousands of specimens from Europe, Asia, and North America, I conclude that Hopkins was correct in treating the North American plants as a different species, A. pycnocarpa. The European A. hirsuta is a diploid (2n = 16) that has fruits 1.5–4 cm, stout styles 0.1–0.5 mm, prominent midvein extending the full length of the fruit valve, and 30–40(–44) ovules/seeds per ovary/fruit. By contrast, A. pycnocarpa is a tetraploid (2n = 32) that has fruits (3.5–)4–6(–6.5) cm, often slender styles (0.2–)0.5–1(–1.3) mm, obscure midvein hardly extending to the middle of the fruit valve, and ovules/seeds (54–)60–86 per ovary/fruit. Hopkins listed other differences in the number of cauline leaves and the development of seed wing, but these do not hold. Mulligan treated the perennial North American plants with minute styles as var. hirsuta and the biennial ones with longer styles as var. pycnocarpa. Habit and style length are not correlated at all, and one finds both biennials and perennials flowering in the same population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 260. | FNA vol. 7, p. 259. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | A. hirsuta var. minshallii, A. pycnocarpa var. reducta | A. hirsuta subsp. pycnocarpa, A. hirsuta var. pycnocarpa | ||||
Name authority | unknown | M. Hopkins: Rhodora 39: 112, plate 458, figs. 1–3. (1937) | ||||
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