Arabis georgiana |
Arabis |
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Georgia rockcress |
rock-cress |
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Habit | Biennials; sparsely to moderately hirsute (at least basally), trichomes simple, mixed with fewer, short-stalked, forked ones, subsessile cruciform or 3-rayed stellate trichomes commonly on abaxial blade surfaces, sometimes plants glabrous distally. | Annuals, biennials, or perennials; (sometimes stoloniferous with vegetative rosettes, or caudex simple or branched); not scapose; often pubescent or hirsute, sometimes glabrous or glabrate, trichomes stalked, stellate, sometimes mixed with fewer, simple or forked ones. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | simple or few from base (rosette), erect, unbranched or branched (few) distally, 3–7 dm, (hirsute basally, glabrous distally). |
erect, ascending, or decumbent, unbranched or branched distally. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate, petiolate or sessile, blade margins usually entire or dentate to denticulate, rarely lyrate-pinnatifid; cauline usually sessile, rarely shortly petiolate, blade (base often auriculate, sagittate, or amplexicaul), margins entire or dentate. |
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Basal leaves | petiole 0.5–2 cm, (ciliate or not); blade spatulate, oblanceolate, or obovate, 1.5–6 cm × 5–15 mm, margins dentate, apex obtuse or acute, abaxial surface moderately to sparsely pubescent, trichomes subsessile stellate, adaxial surface subglabrate or sparsely stellate. |
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Cauline leaves | 7–26; blade oblong, lanceolate, or linear-lanceolate, 1.5–7 cm × 3–18 mm, base auriculate to subamplexicaul, margins dentate or entire, apex acute or obtuse, pubescent as basal leaves except distalmost leaves often glabrous. |
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Racemes | often simple. |
(sometimes paniculate, usually simple, sometimes branched). |
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Flowers | sepals oblong, 2.5–4.5 × 1–1.5 mm, lateral pair subsaccate basally; petals white, narrowly spatulate or oblanceolate, 6–9 × 1–1.5 mm, apex obtuse; filaments 3–4.5 mm; anthers oblong, 0.8–1 mm. |
sepals erect or ascending, ovate or oblong, lateral pair saccate or not basally, (margins membranous); petals white, pink, or purple, usually spatulate, oblong, or oblanceolate, rarely obovate, claw differentiated from blade, (shorter than sepals, apex obtuse or rounded); stamens tetradynamous; filaments usually not dilated basally; anthers ovate, oblong, or linear, (apex obtuse); nectar glands confluent, subtending bases of stamens, lateral glands semiannular or annular, median glands rarely absent, (sometimes toothlike and distinct). |
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Fruiting pedicels | erect to erect-ascending, 7–16 mm, (glabrous). |
erect, ascending, or divaricate, (not reflexed or secund), slender. |
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Fruits | erect to erect-ascending, (often subappressed to rachis), smooth, 4–7 cm × 0.7–0.8 mm; valves each with midvein extending full length or to middle; ovules 38–44 per ovary; style 0.7–1.8 mm. |
siliques, usually sessile, rarely shortly stipitate, linear, smooth or torulose, (usually straight, sometimes slightly curved), flattened, latiseptate; valves (papery), each with obscure or prominent midvein, glabrous; replum (visible), rounded; septum complete, (membranous, translucent, veinless); ovules 10–86[–110] per ovary; style obsolete or distinct; stigma capitate, (sometimes slightly 2-lobed). |
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Seeds | narrowly winged throughout, oblong, 0.9–1.9 × 0.5–0.7 mm; wing to 0.1 mm wide distally. |
uniseriate, flattened, winged or margined, oblong or orbicular; seed coat (smooth or minutely reticulate), not mucilaginous when wetted; cotyledons accumbent. |
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x | = 8. |
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Arabis georgiana |
Arabis |
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Phenology | Flowering Mar–Apr. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Stream banks, roadsides | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0-200 m (0-700 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AL; GA |
North America; Europe; Asia; n, alpine; c Africa; e Africa |
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Discussion | Of conservation concern. Arabis georgiana is most closely related to A. pycnocarpa, from which it is easily distinguished by having narrower fruits, longer petals, and subsessile cruciform or 3-rayed trichomes on abaxial surfaces of basal leaves. It is known only in Alabama from Bibb and Elmore counties and in Georgia from Stewart County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 70 (15 in the flora). As treated by most North American authors (e.g., M. Hopkins 1937; R. C. Rollins 1941, 1993; G. A. Mulligan 1996), Arabis was so broadly delimited that it included species presently assigned to six genera in five tribes (I. A. Al-Shehbaz et al. 2006). These are: Arabidopsis and Turritis (Camelineae), Arabis (Arabideae), Boechera (Boechereae), Pennellia (Halimolobeae), and Streptanthus (Thelypodieae) (see Al-Shehbaz 2003b; M. D. Windham and Al-Shehbaz 2006, 2007). R. C. Rollins (1993) recognized 80 species of Arabis in North America, of which 24 were divided into 40 varieties. By contrast, G. A. Mulligan (1996) recognized 30 species and six varieties in Canada alone. The combination of characteristics that they used to circumscribe the genus (e.g., linear and latiseptate fruits, accumbent cotyledons, and often branched trichomes) evolved repeatedly in Brassicaceae and cannot be relied on in the delimitation of genera (O. Appel and I. A. Al-Shehbaz 2003). The vast majority of North American species have been transferred to Boechera (x = 7), a genus distinct morphologically, cytologically, and molecularly (Al-Shehbaz 2003b; M. D. Windham and Al-Shehbaz 2006, 2007, 2007b). As currently circumscribed, Arabis (x = 8) is a primarily Eurasian genus with only 15 species in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 261. | FNA vol. 7, p. 257. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | R. M. Harper: Torreya 3: 88. (1903) | Linnaeus: Sp. Pl. 2: 664. (1753): Gen. Pl. ed. 5, 298. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |