Arabis blepharophylla |
Arabis |
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coast rock cress, rose rockcress |
rock-cress |
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Habit | Perennials; (caudex simple or branched, covered with persistent petiolar remains); sparsely to moderately pubescent, trichomes simple, forked-stalked, or rarely cruciform or 3-rayed, stalked, stellate. | Annuals, biennials, or perennials; (sometimes stoloniferous with vegetative rosettes, or caudex simple or branched); not scapose; often pubescent or hirsute, sometimes glabrous or glabrate, trichomes stalked, stellate, sometimes mixed with fewer, simple or forked ones. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | simple or few from base (caudex), erect, unbranched or branched (few) distally, 0.6–2.5(–3) dm, (usually pubescent throughout, rarely subglabrate). |
erect, ascending, or decumbent, unbranched or branched distally. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate, petiolate or sessile, blade margins usually entire or dentate to denticulate, rarely lyrate-pinnatifid; cauline usually sessile, rarely shortly petiolate, blade (base often auriculate, sagittate, or amplexicaul), margins entire or dentate. |
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Basal leaves | petiole 0.5–3(–6) cm, (ciliate); blade oblanceolate, spatulate, or obovate, (1.5–)2.1–3.5(–6) cm × (5–)8–20(–25) mm, margins entire or dentate, (ciliate), apex obtuse, surfaces pubescent or glabrous, trichomes simple and forked, sometimes mixed with 3- or 4-rayed stellate ones. |
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Cauline leaves | (2 or) 3–6 (or 7); blade oblong or ovate, 1–2(–4) cm × (2–)4–10(–15) mm, base not auriculate, margins entire or dentate, (ciliate), apex obtuse, surfaces pubescent as basal leaves. |
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Racemes | simple, (dense). |
(sometimes paniculate, usually simple, sometimes branched). |
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Flowers | sepals (purple), oblong, 5–7 × 1.5–2 mm, lateral pair saccate basally; petals purple, spatulate or broadly so, (12–)14–18 × 4–7 mm, apex obtuse or rounded; filaments 6–8 mm; anthers narrowly oblong, 1.2–1.5 mm. |
sepals erect or ascending, ovate or oblong, lateral pair saccate or not basally, (margins membranous); petals white, pink, or purple, usually spatulate, oblong, or oblanceolate, rarely obovate, claw differentiated from blade, (shorter than sepals, apex obtuse or rounded); stamens tetradynamous; filaments usually not dilated basally; anthers ovate, oblong, or linear, (apex obtuse); nectar glands confluent, subtending bases of stamens, lateral glands semiannular or annular, median glands rarely absent, (sometimes toothlike and distinct). |
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Fruiting pedicels | ascending to erect, (3–)5–10(–15) mm. |
erect, ascending, or divaricate, (not reflexed or secund), slender. |
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Fruits | erect or nearly so, slightly torulose, 2–4 cm × 2–3 mm; valves each with prominent midvein extending full length or rarely to middle; ovules 20–28 per ovary; style 0.2–1(–1.5) mm. |
siliques, usually sessile, rarely shortly stipitate, linear, smooth or torulose, (usually straight, sometimes slightly curved), flattened, latiseptate; valves (papery), each with obscure or prominent midvein, glabrous; replum (visible), rounded; septum complete, (membranous, translucent, veinless); ovules 10–86[–110] per ovary; style obsolete or distinct; stigma capitate, (sometimes slightly 2-lobed). |
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Seeds | narrowly winged throughout, orbicular or suborbicular, 2–2.5 mm in diam.; wing 0.2–0.4 mm wide (wider distally). |
uniseriate, flattened, winged or margined, oblong or orbicular; seed coat (smooth or minutely reticulate), not mucilaginous when wetted; cotyledons accumbent. |
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x | = 8. |
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2n | = 16. |
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Arabis blepharophylla |
Arabis |
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Phenology | Flowering Mar–Apr. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Rocky hillsides and bluffs, grassy hillsides, slopes | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 50-200 m (200-700 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA
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North America; Europe; Asia; n, alpine; c Africa; e Africa |
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Discussion | Arabis blepharophylla is an attractive species on the gradual increase in cultivation as an ornamental. It is recorded from Contra Costa, Marin, San Francisco, San Mateo, Santa Cruz, and Sonoma counties; most of the records are based on older collections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 70 (15 in the flora). As treated by most North American authors (e.g., M. Hopkins 1937; R. C. Rollins 1941, 1993; G. A. Mulligan 1996), Arabis was so broadly delimited that it included species presently assigned to six genera in five tribes (I. A. Al-Shehbaz et al. 2006). These are: Arabidopsis and Turritis (Camelineae), Arabis (Arabideae), Boechera (Boechereae), Pennellia (Halimolobeae), and Streptanthus (Thelypodieae) (see Al-Shehbaz 2003b; M. D. Windham and Al-Shehbaz 2006, 2007). R. C. Rollins (1993) recognized 80 species of Arabis in North America, of which 24 were divided into 40 varieties. By contrast, G. A. Mulligan (1996) recognized 30 species and six varieties in Canada alone. The combination of characteristics that they used to circumscribe the genus (e.g., linear and latiseptate fruits, accumbent cotyledons, and often branched trichomes) evolved repeatedly in Brassicaceae and cannot be relied on in the delimitation of genera (O. Appel and I. A. Al-Shehbaz 2003). The vast majority of North American species have been transferred to Boechera (x = 7), a genus distinct morphologically, cytologically, and molecularly (Al-Shehbaz 2003b; M. D. Windham and Al-Shehbaz 2006, 2007, 2007b). As currently circumscribed, Arabis (x = 8) is a primarily Eurasian genus with only 15 species in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 266. | FNA vol. 7, p. 257. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Hooker & Arnott: Bot. Beechey Voy., 321. (1838) | Linnaeus: Sp. Pl. 2: 664. (1753): Gen. Pl. ed. 5, 298. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |