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alpine rockcress, arabis alpina

modest rock-cress, Rogue Canyon rock-cress

Habit Perennials; (stoloniferous, with vegetative rosettes, loosely cespitose to somewhat pulvinate); sparsely to moderately pubescent, trichomes stalked, cruciform, stellate, mixed with simple and forked-stalked ones. Perennials; (caudex usually simple, rarely branched, covered with persistent petiolar remains); sparsely to densely pubescent, trichomes short-stalked, cruci-form, or 3-rayed stellate, (0.3–0.5 mm).
Stems

usually simple from base, erect to ascending, often branched proximally, (0.6–)1–2(–2.5) dm.

simple or few from base (caudex), erect, usually unbranched, rarely branched (few) distally, 2.2–5.5(–6.7) dm, (usually pubescent throughout, rarely subglabrate distally, trichomes stellate).

Basal leaves

petiole 0–1 cm;

blade spatulate, oblanceolate, oblong, or obovate, (0.4–)1–4(–5) cm × (3–)6–15(–20) mm, margins dentate to denticulate, apex obtuse or acute, surfaces usually pubescent, rarely subglabrate, trichomes stellate with simple rays.

petiole 1–4(–5) cm, (rarely minutely ciliate near base);

blade oblanceolate to obovate, (0.8–)2–4.5(–6) cm × (4–)10–20(–25) mm, margins entire, repand, or dentate, (not ciliate), apex obtuse, surfaces pubescent, trichomes short-stalked, stellate.

Cauline leaves

3–5(–6);

blade oblong or ovate, 1–3 cm × 5–15 mm, base subcordate or auriculate, margins usually dentate, rarely subentire, apex acute or obtuse.

(2–)4–6(–9);

blade oblong or ovate, 1–3.5(–4.5) cm × (3–)6–10(–20) mm, base not auriculate, margins entire, repand, or dentate, apex obtuse or acute, surfaces pubescent, trichomes short-stalked, stellate.

Racemes

simple, (lax).

simple, (dense).

Flowers

sepals oblong, 2.5–4(–4.7) × 1–2 mm, lateral pair conspicuously saccate basally;

petals white, spatulate to obovate, 5–8(–9) × 2–3.5 mm, apex obtuse;

filaments 3–5 mm;

anthers oblong, 0.7–1.2 mm.

sepals (purple), oblong, (4–)5–6.5(–8) × 1.5–2 mm, lateral pair saccate basally;

petals purple, spatulate, (10–)12–16(–20) × 4–6(–7) mm, apex rounded;

filaments (4–)5–8 mm;

anthers narrowly oblong, 1.3–2 mm.

Fruiting pedicels

ascending to divaricate, 4–10(–12) mm.

ascending to divaricate, 7–18(–25) mm.

Fruits

ascending to spreading, torulose, 1.7–3.5(–4) cm × 1–1.7 mm;

valves each with midvein absent or obscure, along proximal 1/2;

ovules 34–50 per ovary;

style 0.3–0.6 mm.

suberect to divaricate, sometimes slightly curved, slightly torulose, (2.8–)3.5–6 cm × 1.5–2 mm;

valves each with prominent midvein extending to middle or full length;

ovules 20–34 per ovary;

style 0.5–1(–1.5) mm.

Seeds

narrowly winged throughout, ovate, 1–1.4 × 0.9–1.1 mm;

wing 0.1–0.2 mm wide.

winged distally, oblong, 1.7–2.2 × 0.9–1.2 mm;

wing 0.2–0.5 mm wide.

2n

= 16.

= 32.

Arabis alpina

Arabis modesta

Phenology Flowering Jun–Aug. Flowering Mar–Apr(-May).
Habitat Crevices of limestone rocks, along streams, calcareous alpine meadows, Salix scrub on slopes with scree Moist shaded banks, slopes, rocky canyon walls, talus, basaltic bluffs
Elevation 0-2400 m (0-7900 ft) 150-500 m (500-1600 ft)
Distribution
from FNA
NY; WA; NL; NU; QC; Greenland; Europe; sw Asia; n Africa; tropical Africa
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[BONAP county map]
from FNA
CA; OR
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Discussion

Arabis alpina, the generic type, is variable. It is cultivated extensively for its attractive flowers, and it sometimes escapes from cultivation. There is disagreement as to whether one or more species should be recognized in this complex. European, some African, and most North American plants are recognized as A. alpina; most of the larger-flowered, southwestern Asian plants, which are most commonly cultivated, are recognized as A. caucasica or as A. alpina subsp. caucasica (Willdenow) Briquet. In my opinion, the morphological differences between the two (see key, couplet 3) support recognition of two species, as did R. C. Rollins (1993) and G. A. Mulligan (1996).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arabis modesta, known only from Napa, Siskiyou, and Yolo counties (California), and Jackson and Josephine counties (Oregon), and the following four species form a well-defined group easily separated from the first nine species above by their large, purple petals. Arabis modesta is readily distinguished from other purple-flowered Arabis by having very fine, short-stalked cruciform and 3-rayed, instead of simple and 2-rayed, forked trichomes. Arabis blepharophylla has broadly winged, orbicular seeds, and the other purple-flowered species (A. aculeolata, A. mcdonaldiana, A. oregana) have oblong seeds that are not winged or narrowly winged distally. The lines separating these three species are not as well-defined as those of A. blepharophylla and A. modesta, and further studies are needed to resolve them. I prefer to maintain all five species of purple-flowered Arabis as distinct until thorough molecular and hybridization studies are conducted. In the absence of such studies, it is unwise to make any nomenclatural changes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 259. FNA vol. 7, p. 264.
Parent taxa Brassicaceae > tribe Arabideae > Arabis Brassicaceae > tribe Arabideae > Arabis
Sibling taxa
A. aculeolata, A. blepharophylla, A. caucasica, A. crucisetosa, A. eschscholtziana, A. furcata, A. georgiana, A. mcdonaldiana, A. modesta, A. nuttallii, A. olympica, A. oregana, A. patens, A. pycnocarpa
A. aculeolata, A. alpina, A. blepharophylla, A. caucasica, A. crucisetosa, A. eschscholtziana, A. furcata, A. georgiana, A. mcdonaldiana, A. nuttallii, A. olympica, A. oregana, A. patens, A. pycnocarpa
Synonyms A. alpina var. glabrata
Name authority Linnaeus: Sp. Pl. 2: 664. (1753) Rollins: Rhodora 43: 350. (1941)
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