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Waldo rock-cress

alpine rockcress, arabis alpina

Habit Perennials; (caudex simple or branched, covered with persistent petiolar remains); sparsely to moderately pubescent, trichomes sub-setiform, bulbous-based, simple, (to 1.5 mm), often mixed with fewer, forked, stalked ones. Perennials; (stoloniferous, with vegetative rosettes, loosely cespitose to somewhat pulvinate); sparsely to moderately pubescent, trichomes stalked, cruciform, stellate, mixed with simple and forked-stalked ones.
Stems

simple or few from base (caudex), erect, unbranched, (0.6–)1.5–3.5(–4.5) dm, (sparsely to densely hirsute basally or throughout).

usually simple from base, erect to ascending, often branched proximally, (0.6–)1–2(–2.5) dm.

Basal leaves

petiole 0.3–1.5 cm, (ciliate);

blade oblanceolate to obovate, (0.5–)1–3(–4) cm × (2–)3–6(–10) mm, margins entire, repand, or obtusely dentate, (ciliate), apex obtuse, surfaces pubescent, trichomes usually simple, sometimes mixed with fewer forked ones, rarely subglabrate.

petiole 0–1 cm;

blade spatulate, oblanceolate, oblong, or obovate, (0.4–)1–4(–5) cm × (3–)6–15(–20) mm, margins dentate to denticulate, apex obtuse or acute, surfaces usually pubescent, rarely subglabrate, trichomes stellate with simple rays.

Cauline leaves

3–6 (or 7);

blade oblong, 0.4–1(–1.5) cm × (1–)2–4 mm, base not auriculate, margins entire or repand, apex obtuse.

3–5(–6);

blade oblong or ovate, 1–3 cm × 5–15 mm, base subcordate or auriculate, margins usually dentate, rarely subentire, apex acute or obtuse.

Racemes

simple, (dense).

simple, (lax).

Flowers

sepals (purple), oblong, 4–8 × 1.5–2.5 mm, lateral pair saccate basally;

petals purple, spatulate, (9–)10–18(–20) × (2.5–)3.5–6(–8) mm, apex obtuse;

filaments 4–9 mm;

anthers oblong, 1.5–2 mm.

sepals oblong, 2.5–4(–4.7) × 1–2 mm, lateral pair conspicuously saccate basally;

petals white, spatulate to obovate, 5–8(–9) × 2–3.5 mm, apex obtuse;

filaments 3–5 mm;

anthers oblong, 0.7–1.2 mm.

Fruiting pedicels

ascending to suberect, 8–12(–15) mm, (subglabrate or sparsely pubescent).

ascending to divaricate, 4–10(–12) mm.

Fruits

ascending to suberect, not torulose, sometimes slightly curved, 3.5–6.5 cm × 1.5–2 mm;

valves each with prominent midvein extending full length;

ovules 24–36 per ovary;

style (0.7–)1–2 mm.

ascending to spreading, torulose, 1.7–3.5(–4) cm × 1–1.7 mm;

valves each with midvein absent or obscure, along proximal 1/2;

ovules 34–50 per ovary;

style 0.3–0.6 mm.

Seeds

narrowly winged nearly throughout except wider distally, oblong, 1.5–2.3 × 1–1.3 mm;

wing 0.1–0.3 mm wide.

narrowly winged throughout, ovate, 1–1.4 × 0.9–1.1 mm;

wing 0.1–0.2 mm wide.

2n

= 32.

= 16.

Arabis aculeolata

Arabis alpina

Phenology Flowering Mar–May(-Jun). Flowering Jun–Aug.
Habitat Serpentine slopes, rocky hillsides Crevices of limestone rocks, along streams, calcareous alpine meadows, Salix scrub on slopes with scree
Elevation 200-1800 m (700-5900 ft) 0-2400 m (0-7900 ft)
Distribution
from FNA
OR
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from FNA
NY; WA; NL; NU; QC; Greenland; Europe; sw Asia; n Africa; tropical Africa
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Discussion

Arabis aculeolata is known only from Curry, Douglas, and Josephine counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arabis alpina, the generic type, is variable. It is cultivated extensively for its attractive flowers, and it sometimes escapes from cultivation. There is disagreement as to whether one or more species should be recognized in this complex. European, some African, and most North American plants are recognized as A. alpina; most of the larger-flowered, southwestern Asian plants, which are most commonly cultivated, are recognized as A. caucasica or as A. alpina subsp. caucasica (Willdenow) Briquet. In my opinion, the morphological differences between the two (see key, couplet 3) support recognition of two species, as did R. C. Rollins (1993) and G. A. Mulligan (1996).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 265. FNA vol. 7, p. 259.
Parent taxa Brassicaceae > tribe Arabideae > Arabis Brassicaceae > tribe Arabideae > Arabis
Sibling taxa
A. alpina, A. blepharophylla, A. caucasica, A. crucisetosa, A. eschscholtziana, A. furcata, A. georgiana, A. mcdonaldiana, A. modesta, A. nuttallii, A. olympica, A. oregana, A. patens, A. pycnocarpa
A. aculeolata, A. blepharophylla, A. caucasica, A. crucisetosa, A. eschscholtziana, A. furcata, A. georgiana, A. mcdonaldiana, A. modesta, A. nuttallii, A. olympica, A. oregana, A. patens, A. pycnocarpa
Synonyms A. alpina var. glabrata
Name authority Greene: Leafl. Bot. Observ. Crit. 2: 69. (1910) Linnaeus: Sp. Pl. 2: 664. (1753)
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