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aphragmus

Aleutian cress, Eschscholtz's aphragmus, Eschscholtz's little nightmare

Habit Plants not scapose; (sometimes rhizomatous, caudex thick, covered with persistent petiolar remains); glabrous or pubescent, trichomes simple or forked, (less than 0.1 mm).
Stems

erect or ascending, branched basally.

2–6 cm; glabrous or sparsely puberulent, trichomes simple.

Leaves

basal and cauline;

petiolate or sessile;

basal rosulate, petiolate, blade margins entire;

cauline petiolate or sessile, blade margins entire.

Basal leaves

petiole 0.5–3 cm (base persistent, papery, broadly expanded, narrowly triangular, 0.2–0.4 cm wide);

blade (subfleshy), spatulate, oblanceolate, or ovate, 0.5–1.2 cm × 2–4 mm, base cuneate to attenuate, apex obtuse, surfaces glabrous or puberulent.

Cauline leaves

(bracts), sessile;

blade similar to basal, narrower, smaller distally.

Racemes

(few- to several-flowered), bracteate throughout or basally.

subumbellate.

Flowers

sepals (sometimes persistent), oblong;

petals broadly obovate or spatulate, claw subequaling sepal, (apex rounded [obtuse]);

stamens slightly tetradynamous;

filaments dilated or not basally;

anthers obtuse;

nectar glands confluent, subtending bases of stamens.

sepals often purplish, 1.5–2 × 0.7–1 mm, glabrous;

petals purple with darker veins, 2–3 × 1–2 mm;

filaments 1.5–2 mm;

anthers 0.2–0.4 mm;

gynophore 0–0.7 mm.

Fruiting pedicels

[erect] ascending, divaricate, sometimes recurved, slender.

1.5–5 mm, glabrous or sparsely puberulent adaxially.

Fruits

silicles or siliques, sessile or subsessile, oblong, elliptic, or lanceolate [ovate, linear];

valves each with obscure or distinct midvein, (smooth), glabrous;

replum flattened basally;

septum absent [complete and membranous];

style obsolete or distinct (to 2 mm);

stigma capitate.

compressed, 5–15 × 2–3 mm;

style 0.2–1 mm.

Seeds

plump, not winged, oblong or ovoid (on filiform funicles often longer than seeds);

seed coat (minutely reticulate), not mucilaginous when wetted;

cotyledons incumbent.

light brown, 1.2–1.7 × 0.8–1 mm;

funicles 2–3 mm.

x

= 7.

2n

= 14.

Aphragmus

Aphragmus eschscholtzianus

Phenology Flowering Jun–Jul.
Habitat Limestone slopes, subalpine valleys, wet cliffs, ridges, fine gravel saturated by snowmelt
Elevation 0-2100 m (0-6900 ft)
Distribution
from USDA
nw North America; Asia (Himalaya, Russian Far East)
[BONAP county map]
from FNA
AK; YT; Asia (Russian Far East [Chukchi Peninsula])
[BONAP county map]
Discussion

Species 11 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 256. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 256.
Parent taxa Brassicaceae > tribe Aphragmeae Brassicaceae > tribe Aphragmeae > Aphragmus
Subordinate taxa
A. eschscholtzianus
Synonyms Oreas, Orobium, Staintoniella Eutrema eschscholtzianum
Name authority Andrzejowski ex de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 209. (1824) Andrzejowski ex de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 210. (1824)
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