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brown pussytoes, brown-bract pussytoes, umber or brown or brown-bract pussytoes, umber pussytoes

beautiful pussy-toes, Sierra pussytoes

Habit Dioecious. Dioecious.
Plants

7–16 cm (bases somewhat woody).

(1–)3–12 cm (stems usually stipitate-glandular).

Stolons

7–16 cm (usually erect, slightly woody).

1–4(–9) cm.

Basal leaves

1-nerved, narrowly spatulate to cuneate, 10–17 × 2–5.4 mm, tips mucronate, faces gray-tomentose.

1-nerved, spatulate to linear-cuneate, 6–12 × 1.5–4.5 mm, tips mucronate, faces glabrescent-scabrous to gray-pubescent (often with purple glandular hairs).

Cauline leaves

linear, 8–18 mm, not flagged (apices acute).

linear, 3–11(–13) mm, usually not flagged (apices acute to acuminate), rarely distal flagged.

Involucres

staminate 3–6 mm; pistillate 4–6.5 mm.

staminate 4–5 mm; pistillate 3.5–4.5 mm.

Corollas

staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm.

staminate 1.9–2.8 mm; pistillate 2–3 mm.

Phyllaries

distally whitish, yellowish, or pale brownish (often streaked with pink or rose).

(relatively wide) distally dark brown-black (sometimes light brown or whitish at very tips; apices blunt).

Heads

3–8 in corymbiform arrays.

4–6 in corymbiform arrays.

Cypselae

0.5–1.2 mm, glabrous;

pappi: staminate 3–4.5 mm; pistillate 3–5 mm.

0.7–1.3 mm, glabrous or slightly papillate;

pappi: staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm.

2n

= 28, 56.

= 28 (as A. media).

Antennaria umbrinella

Antennaria pulchella

Phenology Flowering summer. Flowering summer.
Habitat Sagebrush steppe to open, dry, coniferous montane forests to subalpine meadows Moist subalpine-alpine meadows, snow basins, margins of tarns, streams, or run-off from snow masses
Elevation 1100–3400 m (3600–11200 ft) 2800–3700 m (9200–12100 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria umbrinella is a primary sexual progenitor of the A. rosea complex (R. J. Bayer 1990b). It is characterized by somewhat erect, slightly woody stolons and phyllaries that are usually various shades of brown, sometimes white, or streaked with pink or rose (Bayer 1987b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Antennaria pulchella is the diploid progenitor of A. media and, consequently, a progenitor of the A. alpina complex (R. J. Bayer 1990d). The A. rosea and A. parvifolia complexes also have the genome of A. pulchella, shown in the high elevation clones with dark phyllaries in these two polyploid complexes. Antennaria pulchella is differentiated from A. media by shorter pistillate or staminate corollas and shorter cauline leaves (Bayer). This sexually reproducing diploid ranges from the area around Lake Tahoe to the Mt. Whitney region (Bayer).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 408. FNA vol. 19, p. 410.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Synonyms A. aizoides, A. flavescens, A. reflexa A. alpina var. scabra, A. media subsp. ciliata, A. media subsp. pulchella, A. scabra
Name authority Rydberg: Bull. Torrey Bot. Club 24: 302. (1897) Greene: Leafl. Bot. Observ. Crit. 2: 149. (1911)
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