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brown pussytoes, brown-bract pussytoes, umber or brown or brown-bract pussytoes, umber pussytoes

Parlin's pussytoes

Habit Dioecious. Dioecious or gynoecious (staminate plants in equal frequencies as pistillates or none in populations, respectively).
Plants

7–16 cm (bases somewhat woody).

12–35(–45) cm.

Stolons

7–16 cm (usually erect, slightly woody).

3.5–11(–14) cm (mostly decumbent when young).

Basal leaves

1-nerved, narrowly spatulate to cuneate, 10–17 × 2–5.4 mm, tips mucronate, faces gray-tomentose.

3–5-nerved, obovate-spatulate, obovate, rhombic-obovate, or suborbiculate, 30–95 × 12–45 mm, tips mucronate, faces gray-pubescent to floccose-glabrescent.

Cauline leaves

linear, 8–18 mm, not flagged (apices acute).

oblong-lanceolate, 3.5–45 mm, distalmost flagged.

Involucres

staminate 3–6 mm; pistillate 4–6.5 mm.

staminate 6–9 mm; pistillate (7–)8–13 mm.

Corollas

staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm.

staminate 3.5–5 mm; pistillate 4–7 mm.

Phyllaries

distally whitish, yellowish, or pale brownish (often streaked with pink or rose).

distally white.

Heads

3–8 in corymbiform arrays.

4–12(–15) in tight corymbiform arrays.

Cypselae

0.5–1.2 mm, glabrous;

pappi: staminate 3–4.5 mm; pistillate 3–5 mm.

1–2 mm, minutely papillate;

pappi: staminate 4–5 mm; pistillate 5–8 mm.

2n

= 28, 56.

= 56, 84, 70, 112.

Antennaria umbrinella

Antennaria parlinii

Phenology Flowering summer.
Habitat Sagebrush steppe to open, dry, coniferous montane forests to subalpine meadows
Elevation 1100–3400 m (3600–11200 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; MB; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria umbrinella is a primary sexual progenitor of the A. rosea complex (R. J. Bayer 1990b). It is characterized by somewhat erect, slightly woody stolons and phyllaries that are usually various shades of brown, sometimes white, or streaked with pink or rose (Bayer 1987b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

The Antennaria parlinii complex consists of two fairly distinct subspecies that differ in induments of basal leaves (tomentose in subsp. fallax; glabrous in subsp. parlinii) and other characters (R. J. Bayer and G. L. Stebbins 1982). Antennaria parlinii is the most common eastern North American species (Bayer and Stebbins 1982, 1983). This complex of polyploid sexual and apomictic populations is the result of multiple hybridizations among sexual diploid species including A. plantaginifolia, A. racemosa, and A. solitaria (Bayer 1985b; Bayer and D. J. Crawford 1986). A. Cronquist (1945; H. A. Gleason and Cronquist 1991) included A. parlinii within his circumscription of A. plantaginifolia. By not including the hybrid polyploiid within the circumscription of a single one of its sexual progenitors, the circumscription here better portrays the evolutionary relationships between A. parlinii and its sexual progenitors.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually glandless; basal leaves adaxially tomentose
subsp. fallax
1. Stems usually with purple glandular hairs (at least near summits of young flowering stems); basal leaves adaxially green-glabrous
subsp. parlinii
Source FNA vol. 19, p. 408. FNA vol. 19, p. 402.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. parlinii subsp. fallax, A. parlinii subsp. parlinii
Synonyms A. aizoides, A. flavescens, A. reflexa
Name authority Rydberg: Bull. Torrey Bot. Club 24: 302. (1897) Fernald: Gard. & Forest 10: 284. (1897)
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