Antennaria umbrinella |
Antennaria monocephala |
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brown pussytoes, brown-bract pussytoes, umber or brown or brown-bract pussytoes, umber pussytoes |
one-head pussytoes, pygmy pussytoes, single-head pussytoes |
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Habit | Dioecious. | Dioecious or gynoecious (staminates uncommon or in equal frequencies as pistillates, respectively). | ||||
Plants | 7–16 cm (bases somewhat woody). |
5–13 cm (stems usually stipitate-glandular). |
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Stolons | 7–16 cm (usually erect, slightly woody). |
2–4 cm. |
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Basal leaves | 1-nerved, narrowly spatulate to cuneate, 10–17 × 2–5.4 mm, tips mucronate, faces gray-tomentose. |
1-nerved, spatulate to narrowly spatulate or oblanceolate, 9–18 × 2–4 mm, tips mucronate, abaxial faces tomentose, adaxial glabrous or green-glabrescent, or both gray-pubescent. |
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Cauline leaves | linear, 8–18 mm, not flagged (apices acute). |
linear, 4–11 mm, flagged. |
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Involucres | staminate 3–6 mm; pistillate 4–6.5 mm. |
staminate 5–7 mm; pistillate 5–8 mm. |
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Corollas | staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm. |
staminate 2.5–3.5 mm; pistillate 3.5–4 mm. |
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Phyllaries | distally whitish, yellowish, or pale brownish (often streaked with pink or rose). |
distally brown, dark brown, black, or olivaceous. |
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Heads | 3–8 in corymbiform arrays. |
usually borne singly (rarely 2–3). |
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Cypselae | 0.5–1.2 mm, glabrous; pappi: staminate 3–4.5 mm; pistillate 3–5 mm. |
1–1.3 mm, usually glabrous; pappi: staminate 3–4 mm (none in gynoecious populations); pistillate 4–5 mm. |
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2n | = 28, 56. |
= 28, 56, 60?, 70. |
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Antennaria umbrinella |
Antennaria monocephala |
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Phenology | Flowering summer. | |||||
Habitat | Sagebrush steppe to open, dry, coniferous montane forests to subalpine meadows | |||||
Elevation | 1100–3400 m (3600–11200 ft) | |||||
Distribution |
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; SK
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AK; MT; WY; AB; BC; NL; NT; NU; QC; YT; Russian Far East (Chukotka Peninsula)
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Discussion | Antennaria umbrinella is a primary sexual progenitor of the A. rosea complex (R. J. Bayer 1990b). It is characterized by somewhat erect, slightly woody stolons and phyllaries that are usually various shades of brown, sometimes white, or streaked with pink or rose (Bayer 1987b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). It seems reasonable to follow in part E. Hultén’s (1968) broad concept of Antennaria monocephala (R. J. Bayer 1991). Hultén circumscribed it as containing three subspecies. The sexual phase of A. monocephala (i.e., subsp. monocephala and subsp. philonipha) is known from southern Alaska, south of the Brooks Range, and to Yukon Territory and adjacent areas of the Northwest Territories and across the Bering Strait on the Chukotka Peninsula. Within his concept of A. monocephala, Hultén also circumscribed the presumably autopolyploid apomictic form of the species as A. monocephala subsp. angustata, thereby extending the range of the species across the Canadian arctic into Greenland and down the western Cordillera into Montana and Wyoming. Antennaria monocephala subsp. monocephala is an amphimictic progenitor of the A. alpina agamic complex, as well as the sexual progenitor of the apomicts of subsp. angustata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 408. | FNA vol. 19, p. 411. | ||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | A. aizoides, A. flavescens, A. reflexa | A. alpina var. monocephala | ||||
Name authority | Rydberg: Bull. Torrey Bot. Club 24: 302. (1897) | de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 269. (1838) | ||||
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