Antennaria umbrinella
Antennaria howellii
brown pussytoes, brown-bract pussytoes, umber or brown or brown-bract pussytoes, umber pussytoes
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes
7–16 cm (bases somewhat woody).
(6–)8–35 cm (stems sometimes stipitate-glandular).
7–16 cm (usually erect, slightly woody).
1–9(–12) cm.
1-nerved, narrowly spatulate to cuneate, 10–17 × 2–5.4 mm, tips mucronate, faces gray-tomentose.
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.
linear, 8–18 mm, not flagged (apices acute).
linear, 8–40 mm, distal sometimes flagged (apices acute).
staminate 3–6 mm; pistillate 4–6.5 mm.
staminate 6–6.5 mm; pistillate 6–11 mm.
staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm.
staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.
distally whitish, yellowish, or pale brownish (often streaked with pink or rose).
(bases sometimes rose) distally white, cream, or light brown.
3–8 in corymbiform arrays.
3–15 in corymbiform arrays.
0.5–1.2 mm, glabrous;
pappi: staminate 3–4.5 mm; pistillate 3–5 mm.
0.8–2 mm, ± papillate;
pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.
= 28, 56.
= 56, 84, 140 (under A. neodioica).
Antennaria umbrinella
Antennaria howellii
Antennaria umbrinella is a primary sexual progenitor of the A. rosea complex (R. J. Bayer 1990b). It is characterized by somewhat erect, slightly woody stolons and phyllaries that are usually various shades of brown, sometimes white, or streaked with pink or rose (Bayer 1987b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 4 (4 in the flora).
The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal leaves green-glabrous adaxially | → 2 |
1. Basal leaves pubescent adaxially (sometimes glabrescent with age) | → 3 |
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream | subsp. canadensis |
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white | subsp. howellii |
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends) | subsp. neodioica |
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends) | subsp. petaloidea |
- Local floras:
BC, 
CA, 
OR, 
WA - Local Web sites: CalFlora, CalPhotos,
Flora NW, 
PNW Herbaria, Turner Photog. 
WildflowerSearch
iNaturalist (observations)
USDA Plants Database- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
- Local floras: BC, OR, WA
- Local Web sites: CalFlora, CalPhotos, Flora NW,
Go Botany, 
IL Wildflowers, 
KS Wildflowers, 
MD Biodiversity, 
MI Flora, 
MN Wildflowers, PNW Herbaria, Turner Photog. - WildflowerSearch
- iNaturalist (observations)
- USDA Plants Database
- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
