Antennaria umbrinella
Antennaria friesiana
brown pussytoes, brown-bract pussytoes, umber or brown or brown-bract pussytoes, umber pussytoes
Antennaire de Fries, Fries' pussytoes
7–16 cm (bases somewhat woody).
7–15 cm (stems stipitate-glandular, hairs purple).
7–16 cm (usually erect, slightly woody).
0.1–4 cm.
1-nerved, narrowly spatulate to cuneate, 10–17 × 2–5.4 mm, tips mucronate, faces gray-tomentose.
1-nerved, narrowly spatulate to oblanceolate, 11–30 × 2–4 mm, tips mucronate, abaxial faces tomentose, adaxial green-glabrescent to gray-pubescent.
linear, 8–18 mm, not flagged (apices acute).
linear, 4–20 mm, flagged.
staminate 3–6 mm; pistillate 4–6.5 mm.
staminate 4–6.5 mm; pistillate 5.5–8 mm.
staminate 2.5–3.5 mm; pistillate 2.5–3.5 mm.
staminate 2.5–3 mm; pistillate 3–4.5 mm.
distally whitish, yellowish, or pale brownish (often streaked with pink or rose).
distally usually black, light brown, dark brown, or olivaceous, sometimes.
3–8 in corymbiform arrays.
2–6 in corymbiform arrays.
0.5–1.2 mm, glabrous;
pappi: staminate 3–4.5 mm; pistillate 3–5 mm.
1.2–2 mm, glabrous or slightly papillate;
pappi: staminate 3–4 mm; pistillate 3.5–5 mm.
= 28, 56.
= 28, 56, 63, 100+.
Antennaria umbrinella
Antennaria friesiana
Antennaria umbrinella is a primary sexual progenitor of the A. rosea complex (R. J. Bayer 1990b). It is characterized by somewhat erect, slightly woody stolons and phyllaries that are usually various shades of brown, sometimes white, or streaked with pink or rose (Bayer 1987b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 3 (3 in the flora).
The Antennaria friesiana complex consists of subsp. alaskana, subsp. neoalaskana, and subsp. friesiana, the former two are dioecious (sexual) phases of the latter gynoecious (asexual) form. The sexual populations are known from Alaska and cordilleran areas of northern Yukon and adjacent Northwest Territories (R. J. Bayer 1991). The apomictic phase is almost circumpolar, occurring from the central and eastern Siberian plateau eastward across the North American arctic to Greenland (Bayer). E. Hultén (1968) circumscribed a fourth subspecies, A. friesiana subsp. compacta. After studying its morphology, in the field and herbarium, it is apparent that Hultén’s taxon contains at least three incongruous entities that are probably not at all related to the other two subspecies of A. friesiana. Hultén’s subsp. compacta included A. densifolia, which is recognized as a distinct species, and A. crymophila and A. neoalaskana as taxonomic synonyms. Antennaria compacta in the strict sense and A. crymophila are perhaps hybrid apomicts and are treated here in A. alpina (see Bayer for details). Antennaria neoalaskana is treated here as a subspecies of A. friesiana.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Stolons 1–4 cm (usually prostrate, sometimes ascending, usually rooting at tips, plants forming mats); involucres: pistillate 7–8 mm | subsp. neoalaskana |
1. Stolons 0.5–1 cm (erect, usually not rooting at tips, plants not forming mats); involucres: pistillate 5.5–7 mm | → 2 |
2. Plants gynoecious (staminates unknown) | subsp. friesiana |
2. Plants dioecious (staminates and pistillates in equal frequencies in populations) | subsp. alaskana |
- Local floras:
BC, 
CA, 
OR, 
WA - Local Web sites: CalFlora, CalPhotos,
Flora NW, 
PNW Herbaria, Turner Photog. 
WildflowerSearch
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USDA Plants Database- LBJ Wildflower Center
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