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evergreen everlasting, evergreen pussytoes, everlasting pussytoes, shrubby pussytoes, Siskiyou everlasting

one-head pussytoes, pygmy pussytoes, single-head pussytoes

Habit Dioecious. Dioecious or gynoecious (staminates uncommon or in equal frequencies as pistillates, respectively).
Plants

5–12 cm (densely tufted, bases woody; root crowns relatively slender).

5–13 cm (stems usually stipitate-glandular).

Stolons

none.

2–4 cm.

Basal leaves

absent at flowering.

1-nerved, spatulate to narrowly spatulate or oblanceolate, 9–18 × 2–4 mm, tips mucronate, abaxial faces tomentose, adaxial glabrous or green-glabrescent, or both gray-pubescent.

Cauline leaves

spatulate, 5–12 × 2–4 mm, not flagged (apices emarginate or obtuse, abaxial faces tomentose, adaxial green).

linear, 4–11 mm, flagged.

Involucres

staminate 5–9 mm; pistillate 10–15 mm.

staminate 5–7 mm; pistillate 5–8 mm.

Corollas

staminate 4–5 mm; pistillate 5–8 mm.

staminate 2.5–3.5 mm; pistillate 3.5–4 mm.

Phyllaries

(relatively wide) distally white.

distally brown, dark brown, black, or olivaceous.

Heads

borne singly.

usually borne singly (rarely 2–3).

Cypselae

1–2 mm, papillate;

pappi: staminate 4.5–5.5 mm; pistillate 7–9 mm.

1–1.3 mm, usually glabrous;

pappi: staminate 3–4 mm (none in gynoecious populations); pistillate 4–5 mm.

2n

= 28.

= 28, 56, 60?, 70.

Antennaria suffrutescens

Antennaria monocephala

Phenology Flowering early summer.
Habitat Dry, open coniferous woods or barren slopes on serpentine
Elevation 500–1600 m (1600–5200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MT; WY; AB; BC; NL; NT; NU; QC; YT; Russian Far East (Chukotka Peninsula)
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria suffrutescens is characterized by suffrutescent growth form, relatively small, emarginate, adaxially glabrous, coriaceous leaves, and relatively large heads borne singly. It is known only from serpentine soils in open montane pine forests in Curry and Josephine counties, Oregon, and neighboring Del Norte and Humboldt counties, California (R. J. Bayer and G. L. Stebbins 1987). Antennaria suffrutescens may have contributed to the origin of some of the clones of the A. rosea complex (e.g., J. T. Howell 27718, NY).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

It seems reasonable to follow in part E. Hultén’s (1968) broad concept of Antennaria monocephala (R. J. Bayer 1991). Hultén circumscribed it as containing three subspecies. The sexual phase of A. monocephala (i.e., subsp. monocephala and subsp. philonipha) is known from southern Alaska, south of the Brooks Range, and to Yukon Territory and adjacent areas of the Northwest Territories and across the Bering Strait on the Chukotka Peninsula. Within his concept of A. monocephala, Hultén also circumscribed the presumably autopolyploid apomictic form of the species as A. monocephala subsp. angustata, thereby extending the range of the species across the Canadian arctic into Greenland and down the western Cordillera into Montana and Wyoming.

Antennaria monocephala subsp. monocephala is an amphimictic progenitor of the A. alpina agamic complex, as well as the sexual progenitor of the apomicts of subsp. angustata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants gynoecious (staminate plants unknown)
subsp. angustata
1. Plants dioecious (staminates and pistillates in equal frequencies in populations)
subsp. monocephala
Source FNA vol. 19, p. 408. FNA vol. 19, p. 411.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. monocephala subsp. angustata, A. monocephala subsp. monocephala
Synonyms A. alpina var. monocephala
Name authority Greene: Pittonia 3: 277. (1898) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 269. (1838)
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