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evergreen everlasting, evergreen pussytoes, everlasting pussytoes, shrubby pussytoes, Siskiyou everlasting

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Dioecious. Gynoecious (staminate plants very uncommon).
Plants

5–12 cm (densely tufted, bases woody; root crowns relatively slender).

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

none.

1–9(–12) cm.

Basal leaves

absent at flowering.

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

spatulate, 5–12 × 2–4 mm, not flagged (apices emarginate or obtuse, abaxial faces tomentose, adaxial green).

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate 5–9 mm; pistillate 10–15 mm.

staminate 6–6.5 mm; pistillate 6–11 mm.

Corollas

staminate 4–5 mm; pistillate 5–8 mm.

staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.

Phyllaries

(relatively wide) distally white.

(bases sometimes rose) distally white, cream, or light brown.

Heads

borne singly.

3–15 in corymbiform arrays.

Cypselae

1–2 mm, papillate;

pappi: staminate 4.5–5.5 mm; pistillate 7–9 mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.

2n

= 28.

= 56, 84, 140 (under A. neodioica).

Antennaria suffrutescens

Antennaria howellii

Phenology Flowering early summer.
Habitat Dry, open coniferous woods or barren slopes on serpentine
Elevation 500–1600 m (1600–5200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria suffrutescens is characterized by suffrutescent growth form, relatively small, emarginate, adaxially glabrous, coriaceous leaves, and relatively large heads borne singly. It is known only from serpentine soils in open montane pine forests in Curry and Josephine counties, Oregon, and neighboring Del Norte and Humboldt counties, California (R. J. Bayer and G. L. Stebbins 1987). Antennaria suffrutescens may have contributed to the origin of some of the clones of the A. rosea complex (e.g., J. T. Howell 27718, NY).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 408. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Greene: Pittonia 3: 277. (1898) Greene: Pittonia 3: 174. (1897)
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