Antennaria suffrutescens |
Antennaria howellii |
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evergreen everlasting, evergreen pussytoes, everlasting pussytoes, shrubby pussytoes, Siskiyou everlasting |
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes |
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Habit | Dioecious. | Gynoecious (staminate plants very uncommon). | ||||||||||||
Plants | 5–12 cm (densely tufted, bases woody; root crowns relatively slender). |
(6–)8–35 cm (stems sometimes stipitate-glandular). |
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Stolons | none. |
1–9(–12) cm. |
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Basal leaves | absent at flowering. |
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent. |
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Cauline leaves | spatulate, 5–12 × 2–4 mm, not flagged (apices emarginate or obtuse, abaxial faces tomentose, adaxial green). |
linear, 8–40 mm, distal sometimes flagged (apices acute). |
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Involucres | staminate 5–9 mm; pistillate 10–15 mm. |
staminate 6–6.5 mm; pistillate 6–11 mm. |
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Corollas | staminate 4–5 mm; pistillate 5–8 mm. |
staminate 3–4 mm; pistillate 3.5–6.5(–8) mm. |
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Phyllaries | (relatively wide) distally white. |
(bases sometimes rose) distally white, cream, or light brown. |
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Heads | borne singly. |
3–15 in corymbiform arrays. |
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Cypselae | 1–2 mm, papillate; pappi: staminate 4.5–5.5 mm; pistillate 7–9 mm. |
0.8–2 mm, ± papillate; pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm. |
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2n | = 28. |
= 56, 84, 140 (under A. neodioica). |
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Antennaria suffrutescens |
Antennaria howellii |
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Phenology | Flowering early summer. | |||||||||||||
Habitat | Dry, open coniferous woods or barren slopes on serpentine | |||||||||||||
Elevation | 500–1600 m (1600–5200 ft) | |||||||||||||
Distribution |
CA; OR
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CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
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Discussion | Antennaria suffrutescens is characterized by suffrutescent growth form, relatively small, emarginate, adaxially glabrous, coriaceous leaves, and relatively large heads borne singly. It is known only from serpentine soils in open montane pine forests in Curry and Josephine counties, Oregon, and neighboring Del Norte and Humboldt counties, California (R. J. Bayer and G. L. Stebbins 1987). Antennaria suffrutescens may have contributed to the origin of some of the clones of the A. rosea complex (e.g., J. T. Howell 27718, NY). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (4 in the flora). The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 408. | FNA vol. 19, p. 403. | ||||||||||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||||||||||
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Synonyms | A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii | |||||||||||||
Name authority | Greene: Pittonia 3: 277. (1898) | Greene: Pittonia 3: 174. (1897) | ||||||||||||
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