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Antennaire rosée, rosy everlasting, rosy pussytoes

alpine pussytoes, dark pussytoes, Rocky Mountain pussytoes

Habit Gynoecious (staminate plants uncommon). Dioecious or gynoecious (staminate plants rare or in equal frequency to pistillates, respectively).
Plants

4–30 cm.

5–13 cm.

Stolons

1–7 cm.

1–4 cm.

Basal leaves

1-nerved, 8–40 × 2–10 mm, spatulate, oblanceolate, or cuneate, tips mucronate, faces usually gray-pubescent, adaxial sometimes green-glabrous.

1-nerved, spatulate to oblanceolate, 6–19 × 2.5–6 mm, tips mucronate, faces gray-pubescent.

Cauline leaves

linear, 6–36 mm, usually not flagged (apices acute to subulate or with lanceolate flags).

linear, 5–20 mm, not flagged (apices acute).

Involucres

staminate unknown; pistillate 4–10 mm.

staminate (3.5–)4.5–6.5 mm; pistillate 4–8 mm.

Corollas

staminate unknown; pistillate 2.5–6 mm.

staminate 2.5–4.5 mm; pistillate 3–4.5 mm.

Phyllaries

distally brown, cream, gray, green, pink, red, white, or yellow (apices acute or erose-obtuse).

distally dark brown, black, or olivaceous.

Heads

3–20 in corymbiform arrays.

2–5(–9) in corymbiform arrays.

Cypselae

0.7–1.8 mm, glabrous or papillate;

pappi: staminate unknown; pistillate 3.5–6.5 mm.

0.6–1.6 mm, glabrous or papillate;

pappi: staminate 2.5–4.5 mm; pistillate 4–5.5 mm.

2n

= 42, 56, (70).

= 56, 98, 112.

Antennaria rosea

Antennaria media

Phenology Flowering summer.
Habitat Dry, rocky to moist alpine tundra
Elevation 1500–3800 m (4900–12500 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; NT; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (4 in the flora).

Antennaria rosea is the most widespread Antennaria of North America, occurring in dry to moist habitats from near sea level to the alpine zone. The A. rosea polyploid agamic complex is one of the more morphologically diverse complexes of North American Antennaria. It occurs from the western cordillera of North America from southern California, Arizona, and New Mexico north to subarctic Alaska and east to Greenland and, disjunctly, in the Canadian maritime provinces, eastern Quebec, and immediately north of and adjacent to Lake Superior (R. J. Bayer et al. 1991). Antennaria chilensis (including A. chilensis var. magellanica) is a Patagonian endemic that morphologically fits within the circumscription of A. rosea and may well be an amphitropical disjunct member of the complex.

Antennaria rosea is taxonomically confusing; it includes agamospermous microspecies that have been recognized as distinct taxonomic species. Morphometric and isozyme analyses have demonstrated that the primary source of morphologic variability in the complex derives from six sexually reproducing progenitors, A. aromatica, A. corymbosa, A. pulchella, A. microphylla, A. racemosa, and A. umbrinella (R. J. Bayer 1989b, 1990b, 1990c). Additionally, three other sexually reproducing species, A. marginata, A. suffrutescens, and A. rosulata, may have contributed to the genetic complexity of the A. rosea complex (Bayer 1990b). Here, four reasonably distinct subspecies are recognized within the complex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Antennaria media ranges from Arizona to Alaska; dioecious and gynoecious populations are encountered (R. J. Bayer and G. L. Stebbins 1987). The dioecious (sexual) populations are restricted primarily to California and Oregon (Bayer et al. 1990). The main distinction between A. media and A. alpina is flags on distal cauline leaves present in A. alpina and mostly absent in A. media (Bayer 1990d). Phyllaries of the pistillate plants in A. alpina tend to be acute; they are blunter in A. media. At some point, it may be preferable to follow W. L. Jepson ([1923–1925]) and some later authors and treat A. media as a subspecies of A. alpina. Antennaria media appears to be an autopolyploid derivative of A. pulchella; genes from A. pulchella may have introgressed into the A. alpina and A. parvifolia complexes indirectly through A. media.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 20–40 mm; phyllaries distally usually green, pink, red or white, seldom brown
subsp. rosea
1. Basal leaves 8–20 mm; phyllaries distally brown, cream, gray, green, pink, red, white, or yellow
→ 2
2. Pistillate: involucres 4–6.5 mm, corollas 2.5–4, pappi 3.5–5; cauline leaves 6–20 mm (tips subulate); phyllaries usually distally brown, sometimes cream, gray, or yellow
subsp. confinis
2. Pistillate: involucres 6.5–10 mm, corollas 3.5–6 mm, pappi 5–6.5 mm; cauline leaves 6–19 or 9–26 mm (tips sometimes with flat, lanceolate scarious appendages); phyllaries distally brown, green, pink, red, or white
→ 3
3. Plants 19–30 cm; cauline leaves 9–26 mm (proximalmost usually 19+ mm); heads usu- ally 6–12
subsp. arida
3. Plants 4–17 cm; cauline leaves 6–19 mm (proximalmost usually less than 19 mm); heads usually 3–5
subsp. pulvinata
Source FNA vol. 19, p. 408. FNA vol. 19, p. 411.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. rosea subsp. arida, A. rosea subsp. confinis, A. rosea subsp. pulvinata, A. rosea subsp. rosea
Synonyms A. alpina var. media, A. austromontana, A. candida, A. densa, A. modesta, A. mucronata
Name authority Greene: Pittonia 3: 281. (1898) Greene: Pittonia 3: 286. (1898)
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